EuRBPDB

Description

RBDs

Transcripts

Gene Model

Pathways

PPI

Orthologs

Description

Ensembl ID: ENSCHIG00000018346 (Gene tree)
Gene ID: 102191810
Gene Symbol: GTPBP4
Alias: CRFG|FLJ10686|FLJ10690|NGB|NOG1
Full Name: GTP binding protein 4
Gene Type: protein_coding
Species: Capra_hircus
Status: putative
Strand: Minus strand
Length: 13371 bases
Position: chr13:45745323-45758693
Accession: 102191810
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
ENSCHIP00000019345	NOG1	PF06858.14	6.9e-26	1	1
ENSCHIP00000019337	NOG1	PF06858.14	7.2e-26	1	1
ENSCHIP00000019316	NOG1	PF06858.14	7.2e-26	1	1
ENSCHIP00000019337	MMR_HSR1	PF01926.23	2.8e-18	1	1
ENSCHIP00000019316	MMR_HSR1	PF01926.23	2.8e-18	1	1
ENSCHIP00000019345	MMR_HSR1	PF01926.23	3e-18	1	1
ENSCHIP00000019316	NOGCT	PF08155.11	1.6e-21	1	1
ENSCHIP00000019345	NOGCT	PF08155.11	3.8e-21	1	1

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
ENSCHIT00000027160	GTPBP4-203	1842	-	ENSCHIP00000019345	613 (aa)	-	UPI000D18FC40
ENSCHIT00000027129	GTPBP4-201	1905	XM_005688151	ENSCHIP00000019316	634 (aa)	XP_005688208	UPI0003AF57D0
ENSCHIT00000027152	GTPBP4-202	1866	-	ENSCHIP00000019337	621 (aa)	-	UPI000D194D02

Gene Model

Click here to download ENSCHIG00000018346's gene model file

Pathways

Pathway ID	Pathway Name	Source
chx03008	Ribosome biogenesis in eukaryotes	KEGG

Protein-Protein Interaction (PPI)

Clik here to download ENSCHIG00000018346's network

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
ENSCHIG00000018346	GTPBP4	100	93.849	ENSG00000107937	GTPBP4	100	99.286	Homo_sapiens
ENSCHIG00000018346	GTPBP4	100	82.650	ENSAPOG00000020433	gtpbp4	100	79.213	Acanthochromis_polyacanthus
ENSCHIG00000018346	GTPBP4	100	92.587	ENSAMEG00000001845	GTPBP4	100	92.587	Ailuropoda_melanoleuca
ENSCHIG00000018346	GTPBP4	100	79.022	ENSACIG00000018629	gtpbp4	100	79.022	Amphilophus_citrinellus
ENSCHIG00000018346	GTPBP4	100	82.808	ENSAOCG00000000427	gtpbp4	93	82.808	Amphiprion_ocellaris
ENSCHIG00000018346	GTPBP4	100	82.965	ENSAPEG00000002544	gtpbp4	100	82.965	Amphiprion_percula
ENSCHIG00000018346	GTPBP4	100	83.438	ENSATEG00000012635	gtpbp4	95	83.438	Anabas_testudineus
ENSCHIG00000018346	GTPBP4	52	80.363	ENSAPLG00000013790	-	100	80.363	Anas_platyrhynchos
ENSCHIG00000018346	GTPBP4	100	84.227	ENSACAG00000002943	GTPBP4	100	84.227	Anolis_carolinensis
ENSCHIG00000018346	GTPBP4	87	86.994	ENSANAG00000029209	-	100	86.994	Aotus_nancymaae
ENSCHIG00000018346	GTPBP4	99	81.646	ENSANAG00000022188	-	99	81.487	Aotus_nancymaae
ENSCHIG00000018346	GTPBP4	73	86.637	ENSACLG00000026330	gtpbp4	92	93.333	Astatotilapia_calliptera
ENSCHIG00000018346	GTPBP4	100	82.492	ENSAMXG00000019771	gtpbp4	100	82.492	Astyanax_mexicanus
ENSCHIG00000018346	GTPBP4	100	99.054	ENSBTAG00000014917	GTPBP4	100	99.054	Bos_taurus
ENSCHIG00000018346	GTPBP4	99	52.624	WBGene00020297	nog-1	99	52.624	Caenorhabditis_elegans
ENSCHIG00000018346	GTPBP4	100	93.553	ENSCJAG00000008494	-	100	93.868	Callithrix_jacchus
ENSCHIG00000018346	GTPBP4	100	92.138	ENSCJAG00000022962	-	100	92.453	Callithrix_jacchus
ENSCHIG00000018346	GTPBP4	93	93.526	ENSCAFG00000005536	GTPBP4	100	93.526	Canis_familiaris
ENSCHIG00000018346	GTPBP4	100	93.849	ENSCAFG00020004307	GTPBP4	100	93.849	Canis_lupus_dingo
ENSCHIG00000018346	GTPBP4	100	92.587	ENSTSYG00000004977	GTPBP4	100	92.587	Carlito_syrichta
ENSCHIG00000018346	GTPBP4	99	81.747	ENSCAPG00000002134	GTPBP4	99	81.747	Cavia_aperea
ENSCHIG00000018346	GTPBP4	100	93.533	ENSCPOG00000013736	GTPBP4	100	93.533	Cavia_porcellus
ENSCHIG00000018346	GTPBP4	100	70.505	ENSCCAG00000030115	-	100	69.854	Cebus_capucinus
ENSCHIG00000018346	GTPBP4	99	93.049	ENSCCAG00000021627	-	92	93.523	Cebus_capucinus
ENSCHIG00000018346	GTPBP4	100	94.164	ENSCATG00000004303	GTPBP4	100	94.164	Cercocebus_atys
ENSCHIG00000018346	GTPBP4	100	94.006	ENSCLAG00000011323	GTPBP4	100	94.006	Chinchilla_lanigera
ENSCHIG00000018346	GTPBP4	100	94.322	ENSCSAG00000014097	GTPBP4	100	94.322	Chlorocebus_sabaeus
ENSCHIG00000018346	GTPBP4	96	95.418	ENSCHOG00000012602	GTPBP4	88	95.418	Choloepus_hoffmanni
ENSCHIG00000018346	GTPBP4	100	86.278	ENSCPBG00000008398	GTPBP4	100	86.278	Chrysemys_picta_bellii
ENSCHIG00000018346	GTPBP4	100	63.411	ENSCING00000001957	-	100	64.031	Ciona_intestinalis
ENSCHIG00000018346	GTPBP4	100	62.058	ENSCSAVG00000009793	-	99	78.317	Ciona_savignyi
ENSCHIG00000018346	GTPBP4	100	89.796	ENSCANG00000014168	GTPBP4	100	89.796	Colobus_angolensis_palliatus
ENSCHIG00000018346	GTPBP4	100	93.533	ENSCGRG00001013765	Gtpbp4	100	93.533	Cricetulus_griseus_chok1gshd
ENSCHIG00000018346	GTPBP4	100	93.533	ENSCGRG00000012171	Gtpbp4	100	93.533	Cricetulus_griseus_crigri
ENSCHIG00000018346	GTPBP4	100	78.233	ENSCSEG00000007352	gtpbp4	100	78.233	Cynoglossus_semilaevis
ENSCHIG00000018346	GTPBP4	100	82.019	ENSCVAG00000008993	gtpbp4	100	82.019	Cyprinodon_variegatus
ENSCHIG00000018346	GTPBP4	100	82.177	ENSDARG00000018961	gtpbp4	100	82.177	Danio_rerio
ENSCHIG00000018346	GTPBP4	67	91.059	ENSDNOG00000006746	GTPBP4	100	91.059	Dasypus_novemcinctus
ENSCHIG00000018346	GTPBP4	82	100.000	ENSDORG00000010884	Gtpbp4	96	100.000	Dipodomys_ordii
ENSCHIG00000018346	GTPBP4	100	59.112	FBgn0028473	Non1	100	59.112	Drosophila_melanogaster
ENSCHIG00000018346	GTPBP4	77	87.755	ENSETEG00000016840	-	71	87.755	Echinops_telfairi
ENSCHIG00000018346	GTPBP4	99	63.681	ENSEBUG00000010311	gtpbp4	99	62.866	Eptatretus_burgeri
ENSCHIG00000018346	GTPBP4	100	94.479	ENSEASG00005019082	GTPBP4	100	94.479	Equus_asinus_asinus
ENSCHIG00000018346	GTPBP4	100	94.479	ENSECAG00000001700	GTPBP4	100	94.479	Equus_caballus
ENSCHIG00000018346	GTPBP4	100	93.531	ENSEEUG00000009889	GTPBP4	91	93.531	Erinaceus_europaeus
ENSCHIG00000018346	GTPBP4	100	81.761	ENSELUG00000008578	gtpbp4	96	82.283	Esox_lucius
ENSCHIG00000018346	GTPBP4	100	94.322	ENSFCAG00000026213	GTPBP4	100	94.322	Felis_catus
ENSCHIG00000018346	GTPBP4	99	85.737	ENSFALG00000006971	GTPBP4	87	86.909	Ficedula_albicollis
ENSCHIG00000018346	GTPBP4	100	93.849	ENSFDAG00000000998	GTPBP4	100	93.849	Fukomys_damarensis
ENSCHIG00000018346	GTPBP4	100	82.965	ENSFHEG00000012922	gtpbp4	100	82.965	Fundulus_heteroclitus
ENSCHIG00000018346	GTPBP4	100	81.230	ENSGMOG00000016090	gtpbp4	100	81.230	Gadus_morhua
ENSCHIG00000018346	GTPBP4	100	86.593	ENSGALG00000006703	GTPBP4	100	86.593	Gallus_gallus
ENSCHIG00000018346	GTPBP4	100	83.438	ENSGAFG00000002075	gtpbp4	100	83.438	Gambusia_affinis
ENSCHIG00000018346	GTPBP4	100	82.019	ENSGACG00000017481	gtpbp4	100	82.019	Gasterosteus_aculeatus
ENSCHIG00000018346	GTPBP4	93	86.031	ENSGAGG00000018689	GTPBP4	100	86.031	Gopherus_agassizii
ENSCHIG00000018346	GTPBP4	100	94.006	ENSGGOG00000013372	GTPBP4	100	94.006	Gorilla_gorilla
ENSCHIG00000018346	GTPBP4	100	83.123	ENSHBUG00000000192	gtpbp4	100	83.123	Haplochromis_burtoni
ENSCHIG00000018346	GTPBP4	100	93.375	ENSHGLG00000013071	GTPBP4	100	93.375	Heterocephalus_glaber_female
ENSCHIG00000018346	GTPBP4	100	93.375	ENSHGLG00100017850	GTPBP4	100	93.375	Heterocephalus_glaber_male
ENSCHIG00000018346	GTPBP4	100	85.512	ENSHGLG00100002964	-	100	85.512	Heterocephalus_glaber_male
ENSCHIG00000018346	GTPBP4	100	83.150	ENSHCOG00000011543	gtpbp4	100	83.150	Hippocampus_comes
ENSCHIG00000018346	GTPBP4	100	83.596	ENSIPUG00000014417	GTPBP4	100	83.596	Ictalurus_punctatus
ENSCHIG00000018346	GTPBP4	100	93.849	ENSSTOG00000010042	-	100	93.849	Ictidomys_tridecemlineatus
ENSCHIG00000018346	GTPBP4	100	84.858	ENSSTOG00000023023	-	100	84.858	Ictidomys_tridecemlineatus
ENSCHIG00000018346	GTPBP4	100	91.956	ENSJJAG00000010021	Gtpbp4	100	91.956	Jaculus_jaculus
ENSCHIG00000018346	GTPBP4	100	81.230	ENSKMAG00000021705	gtpbp4	100	81.230	Kryptolebias_marmoratus
ENSCHIG00000018346	GTPBP4	100	78.425	ENSLBEG00000021419	gtpbp4	100	81.973	Labrus_bergylta
ENSCHIG00000018346	GTPBP4	100	83.754	ENSLACG00000001449	GTPBP4	100	83.754	Latimeria_chalumnae
ENSCHIG00000018346	GTPBP4	100	82.000	ENSLOCG00000012279	gtpbp4	100	82.000	Lepisosteus_oculatus
ENSCHIG00000018346	GTPBP4	100	92.744	ENSLAFG00000018578	GTPBP4	100	92.744	Loxodonta_africana
ENSCHIG00000018346	GTPBP4	100	93.849	ENSMFAG00000046251	GTPBP4	100	93.849	Macaca_fascicularis
ENSCHIG00000018346	GTPBP4	100	91.009	ENSMMUG00000010813	GTPBP4	100	91.009	Macaca_mulatta
ENSCHIG00000018346	GTPBP4	100	94.006	ENSMNEG00000044366	GTPBP4	100	94.006	Macaca_nemestrina
ENSCHIG00000018346	GTPBP4	100	93.849	ENSMLEG00000039605	GTPBP4	100	93.849	Mandrillus_leucophaeus
ENSCHIG00000018346	GTPBP4	100	83.281	ENSMAMG00000013763	gtpbp4	92	83.281	Mastacembelus_armatus
ENSCHIG00000018346	GTPBP4	100	83.281	ENSMZEG00005006621	gtpbp4	100	83.281	Maylandia_zebra
ENSCHIG00000018346	GTPBP4	100	93.218	ENSMAUG00000000571	Gtpbp4	100	93.218	Mesocricetus_auratus
ENSCHIG00000018346	GTPBP4	100	93.375	ENSMICG00000013332	GTPBP4	100	93.375	Microcebus_murinus
ENSCHIG00000018346	GTPBP4	100	93.533	ENSMOCG00000002209	Gtpbp4	100	93.533	Microtus_ochrogaster
ENSCHIG00000018346	GTPBP4	100	79.338	ENSMMOG00000011257	gtpbp4	100	79.338	Mola_mola
ENSCHIG00000018346	GTPBP4	100	90.063	ENSMODG00000005978	GTPBP4	100	90.063	Monodelphis_domestica
ENSCHIG00000018346	GTPBP4	100	81.546	ENSMALG00000012488	gtpbp4	100	81.546	Monopterus_albus
ENSCHIG00000018346	GTPBP4	99	91.614	MGP_CAROLIEiJ_G0018215	Gtpbp4	100	91.667	Mus_caroli
ENSCHIG00000018346	GTPBP4	100	93.060	ENSMUSG00000021149	Gtpbp4	100	93.060	Mus_musculus
ENSCHIG00000018346	GTPBP4	100	92.902	MGP_PahariEiJ_G0019265	Gtpbp4	100	92.902	Mus_pahari
ENSCHIG00000018346	GTPBP4	99	92.880	MGP_SPRETEiJ_G0019073	Gtpbp4	100	92.880	Mus_spretus
ENSCHIG00000018346	GTPBP4	93	93.186	ENSMPUG00000016221	GTPBP4	100	93.186	Mustela_putorius_furo
ENSCHIG00000018346	GTPBP4	100	93.375	ENSMLUG00000028192	GTPBP4	100	93.375	Myotis_lucifugus
ENSCHIG00000018346	GTPBP4	100	92.587	ENSNGAG00000015364	Gtpbp4	100	92.587	Nannospalax_galili
ENSCHIG00000018346	GTPBP4	87	82.222	ENSNBRG00000019965	gtpbp4	86	82.222	Neolamprologus_brichardi
ENSCHIG00000018346	GTPBP4	100	94.006	ENSNLEG00000013495	GTPBP4	100	94.006	Nomascus_leucogenys
ENSCHIG00000018346	GTPBP4	66	90.909	ENSMEUG00000016215	-	72	90.909	Notamacropus_eugenii
ENSCHIG00000018346	GTPBP4	100	95.957	ENSOPRG00000008671	GTPBP4	91	95.957	Ochotona_princeps
ENSCHIG00000018346	GTPBP4	100	94.322	ENSODEG00000000962	-	100	94.322	Octodon_degus
ENSCHIG00000018346	GTPBP4	100	92.114	ENSODEG00000019152	-	100	92.114	Octodon_degus
ENSCHIG00000018346	GTPBP4	100	83.281	ENSONIG00000017707	gtpbp4	100	83.281	Oreochromis_niloticus
ENSCHIG00000018346	GTPBP4	99	87.697	ENSOANG00000006095	-	99	87.697	Ornithorhynchus_anatinus
ENSCHIG00000018346	GTPBP4	61	85.488	ENSOCUG00000014716	-	100	85.488	Oryctolagus_cuniculus
ENSCHIG00000018346	GTPBP4	100	91.009	ENSOCUG00000016120	-	100	91.009	Oryctolagus_cuniculus
ENSCHIG00000018346	GTPBP4	100	82.177	ENSORLG00000006151	gtpbp4	100	82.177	Oryzias_latipes
ENSCHIG00000018346	GTPBP4	100	82.177	ENSORLG00020020899	gtpbp4	100	82.177	Oryzias_latipes_hni
ENSCHIG00000018346	GTPBP4	100	82.019	ENSORLG00015020562	gtpbp4	100	82.019	Oryzias_latipes_hsok
ENSCHIG00000018346	GTPBP4	100	82.019	ENSOMEG00000019684	gtpbp4	100	82.019	Oryzias_melastigma
ENSCHIG00000018346	GTPBP4	100	93.849	ENSOGAG00000011941	GTPBP4	100	93.849	Otolemur_garnettii
ENSCHIG00000018346	GTPBP4	100	98.901	ENSOARG00000015047	GTPBP4	100	98.901	Ovis_aries
ENSCHIG00000018346	GTPBP4	100	94.006	ENSPPAG00000042066	GTPBP4	100	94.006	Pan_paniscus
ENSCHIG00000018346	GTPBP4	100	94.322	ENSPPRG00000009895	GTPBP4	100	94.322	Panthera_pardus
ENSCHIG00000018346	GTPBP4	100	94.164	ENSPTIG00000008569	GTPBP4	100	94.164	Panthera_tigris_altaica
ENSCHIG00000018346	GTPBP4	73	94.667	ENSPTRG00000002236	GTPBP4	86	94.866	Pan_troglodytes
ENSCHIG00000018346	GTPBP4	100	94.006	ENSPANG00000023543	GTPBP4	100	94.006	Papio_anubis
ENSCHIG00000018346	GTPBP4	100	82.650	ENSPKIG00000012301	gtpbp4	100	82.650	Paramormyrops_kingsleyae
ENSCHIG00000018346	GTPBP4	58	91.597	ENSPSIG00000005442	GTPBP4	93	91.597	Pelodiscus_sinensis
ENSCHIG00000018346	GTPBP4	100	79.022	ENSPMGG00000001462	gtpbp4	100	79.022	Periophthalmus_magnuspinnatus
ENSCHIG00000018346	GTPBP4	100	92.744	ENSPEMG00000015091	-	100	92.744	Peromyscus_maniculatus_bairdii
ENSCHIG00000018346	GTPBP4	100	69.381	ENSPEMG00000012279	-	100	69.381	Peromyscus_maniculatus_bairdii
ENSCHIG00000018346	GTPBP4	100	76.025	ENSPMAG00000000905	gtpbp4	100	76.025	Petromyzon_marinus
ENSCHIG00000018346	GTPBP4	100	90.063	ENSPCIG00000026082	GTPBP4	100	90.063	Phascolarctos_cinereus
ENSCHIG00000018346	GTPBP4	100	83.123	ENSPFOG00000000026	gtpbp4	100	83.123	Poecilia_formosa
ENSCHIG00000018346	GTPBP4	92	83.419	ENSPLAG00000002898	gtpbp4	97	83.419	Poecilia_latipinna
ENSCHIG00000018346	GTPBP4	100	83.281	ENSPMEG00000016422	gtpbp4	100	83.281	Poecilia_mexicana
ENSCHIG00000018346	GTPBP4	100	79.811	ENSPREG00000019870	gtpbp4	100	79.811	Poecilia_reticulata
ENSCHIG00000018346	GTPBP4	100	94.322	ENSPPYG00000002037	GTPBP4	100	94.322	Pongo_abelii
ENSCHIG00000018346	GTPBP4	53	93.333	ENSPCAG00000006152	-	50	93.333	Procavia_capensis
ENSCHIG00000018346	GTPBP4	97	90.285	ENSPCOG00000014602	-	100	90.285	Propithecus_coquereli
ENSCHIG00000018346	GTPBP4	100	88.486	ENSPCOG00000014382	-	100	88.486	Propithecus_coquereli
ENSCHIG00000018346	GTPBP4	92	94.969	ENSPVAG00000005882	GTPBP4	86	94.969	Pteropus_vampyrus
ENSCHIG00000018346	GTPBP4	100	73.817	ENSPNYG00000024251	gtpbp4	100	73.817	Pundamilia_nyererei
ENSCHIG00000018346	GTPBP4	100	83.123	ENSPNAG00000019977	gtpbp4	100	83.123	Pygocentrus_nattereri
ENSCHIG00000018346	GTPBP4	100	92.610	ENSRNOG00000016217	Gtpbp4	100	92.767	Rattus_norvegicus
ENSCHIG00000018346	GTPBP4	100	94.322	ENSRBIG00000028875	GTPBP4	100	94.322	Rhinopithecus_bieti
ENSCHIG00000018346	GTPBP4	100	94.322	ENSRROG00000030798	GTPBP4	100	94.322	Rhinopithecus_roxellana
ENSCHIG00000018346	GTPBP4	99	46.225	YPL093W	-	99	46.225	Saccharomyces_cerevisiae
ENSCHIG00000018346	GTPBP4	100	93.711	ENSSBOG00000028421	GTPBP4	100	93.711	Saimiri_boliviensis_boliviensis
ENSCHIG00000018346	GTPBP4	99	89.968	ENSSHAG00000014713	GTPBP4	94	89.968	Sarcophilus_harrisii
ENSCHIG00000018346	GTPBP4	100	83.281	ENSSFOG00015003167	gtpbp4	100	83.281	Scleropages_formosus
ENSCHIG00000018346	GTPBP4	100	82.334	ENSSMAG00000005863	gtpbp4	100	82.334	Scophthalmus_maximus
ENSCHIG00000018346	GTPBP4	100	82.019	ENSSDUG00000006534	gtpbp4	100	82.019	Seriola_dumerili
ENSCHIG00000018346	GTPBP4	100	82.334	ENSSLDG00000020688	gtpbp4	100	82.334	Seriola_lalandi_dorsalis
ENSCHIG00000018346	GTPBP4	61	99.010	ENSSARG00000003409	-	79	99.010	Sorex_araneus
ENSCHIG00000018346	GTPBP4	100	86.278	ENSSPUG00000002153	GTPBP4	100	86.278	Sphenodon_punctatus
ENSCHIG00000018346	GTPBP4	63	72.040	ENSSPAG00000014583	gtpbp4	100	72.040	Stegastes_partitus
ENSCHIG00000018346	GTPBP4	100	93.060	ENSSSCG00000028602	GTPBP4	100	93.060	Sus_scrofa
ENSCHIG00000018346	GTPBP4	99	85.350	ENSTGUG00000000820	GTPBP4	100	86.306	Taeniopygia_guttata
ENSCHIG00000018346	GTPBP4	100	81.546	ENSTRUG00000003900	gtpbp4	100	81.546	Takifugu_rubripes
ENSCHIG00000018346	GTPBP4	100	83.281	ENSTNIG00000007992	gtpbp4	100	83.281	Tetraodon_nigroviridis
ENSCHIG00000018346	GTPBP4	100	79.935	ENSTBEG00000003613	GTPBP4	100	79.935	Tupaia_belangeri
ENSCHIG00000018346	GTPBP4	100	92.892	ENSTTRG00000011163	GTPBP4	100	92.892	Tursiops_truncatus
ENSCHIG00000018346	GTPBP4	100	93.375	ENSUAMG00000013191	GTPBP4	100	93.375	Ursus_americanus
ENSCHIG00000018346	GTPBP4	93	92.675	ENSUMAG00000016138	GTPBP4	100	92.675	Ursus_maritimus
ENSCHIG00000018346	GTPBP4	100	93.691	ENSVVUG00000008608	GTPBP4	98	93.691	Vulpes_vulpes
ENSCHIG00000018346	GTPBP4	100	73.502	ENSXETG00000018097	gtpbp4	100	74.474	Xenopus_tropicalis
ENSCHIG00000018346	GTPBP4	100	77.080	ENSXCOG00000019953	gtpbp4	98	77.080	Xiphophorus_couchianus
ENSCHIG00000018346	GTPBP4	100	82.575	ENSXMAG00000006719	gtpbp4	100	82.575	Xiphophorus_maculatus