EuRBPDB

Description

RBDs

Transcripts

Gene Model

Pathways

PPI

Orthologs

Description

Ensembl ID: ENSCPBG00000001138 (Gene tree)
Gene ID: 101945149
Gene Symbol: EPRS
Alias: EARS|GLUPRORS|PARS|QARS|QPRS
Full Name: glutamyl-prolyl-tRNA synthetase
Gene Type: protein_coding
Species: Chrysemys_picta_bellii
Status: putative
Strand: Plus strand
Length: 68999 bases
Position: chrKK088190.1:1356202-1425200
Accession: 3418
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
ENSCPBP00000017261	tRNA-synt_1c_C	PF03950.18	1.1e-37	1	1
ENSCPBP00000016620	tRNA-synt_1c_C	PF03950.18	1.2e-37	1	1
ENSCPBP00000016614	tRNA-synt_1c_C	PF03950.18	1.3e-37	1	1
ENSCPBP00000017524	tRNA-synt_1c_C	PF03950.18	1.3e-37	1	1
ENSCPBP00000017296	tRNA-synt_1c_C	PF03950.18	1.6e-37	1	1
ENSCPBP00000017670	tRNA-synt_1c_C	PF03950.18	1.9e-14	1	1
ENSCPBP00000017261	HGTP_anticodon	PF03129.20	3.1e-17	1	1
ENSCPBP00000017670	HGTP_anticodon	PF03129.20	3.4e-17	1	1
ENSCPBP00000017524	HGTP_anticodon	PF03129.20	4e-17	1	1
ENSCPBP00000016620	HGTP_anticodon	PF03129.20	4.3e-17	1	1
ENSCPBP00000016614	HGTP_anticodon	PF03129.20	4.5e-17	1	1
ENSCPBP00000017296	HGTP_anticodon	PF03129.20	4.9e-17	1	1
ENSCPBP00000017670	tRNA-synt_1c	PF00749.21	8.3e-114	1	1
ENSCPBP00000017524	tRNA-synt_1c	PF00749.21	8.8e-114	1	1
ENSCPBP00000017296	tRNA-synt_1c	PF00749.21	1e-113	1	1
ENSCPBP00000016614	tRNA-synt_1c	PF00749.21	1e-113	1	1
ENSCPBP00000016620	tRNA-synt_1c	PF00749.21	8.4e-113	1	1
ENSCPBP00000017261	tRNA-synt_1c	PF00749.21	9.3e-88	1	2
ENSCPBP00000017261	tRNA-synt_1c	PF00749.21	9.3e-88	2	2
ENSCPBP00000016620	tRNA-synt_2b	PF00587.25	1e-16	1	1
ENSCPBP00000017296	tRNA-synt_2b	PF00587.25	1.1e-16	1	1
ENSCPBP00000017524	tRNA-synt_2b	PF00587.25	1.1e-16	1	1
ENSCPBP00000016614	tRNA-synt_2b	PF00587.25	1.1e-16	1	1
ENSCPBP00000017670	tRNA-synt_2b	PF00587.25	1.3e-16	1	1
ENSCPBP00000017261	tRNA-synt_2b	PF00587.25	1.8e-16	1	1
ENSCPBP00000016614	ProRS-C_1	PF09180.11	6.1e-22	1	1
ENSCPBP00000017296	ProRS-C_1	PF09180.11	6.1e-22	1	1
ENSCPBP00000017261	ProRS-C_1	PF09180.11	6.4e-22	1	1
ENSCPBP00000017670	ProRS-C_1	PF09180.11	1.8e-21	1	1
ENSCPBP00000017524	ProRS-C_1	PF09180.11	1.9e-21	1	1
ENSCPBP00000016620	ProRS-C_1	PF09180.11	1.9e-21	1	1

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
ENSCPBT00000020719	EPRS-205	4395	-	ENSCPBP00000017524	1464 (aa)	-	-
ENSCPBT00000019651	EPRS-202	4530	-	ENSCPBP00000016620	1509 (aa)	-	-
ENSCPBT00000019645	EPRS-201	4764	-	ENSCPBP00000016614	1587 (aa)	-	-
ENSCPBT00000020455	EPRS-204	4758	-	ENSCPBP00000017296	1585 (aa)	-	-
ENSCPBT00000020875	EPRS-206	4224	-	ENSCPBP00000017670	1407 (aa)	-	-
ENSCPBT00000020418	EPRS-203	4353	-	ENSCPBP00000017261	1450 (aa)	-	-

Gene Model

Click here to download ENSCPBG00000001138's gene model file

Pathways

Pathway ID	Pathway Name	Source
cpic00860	Porphyrin and chlorophyll metabolism	KEGG
cpic00970	Aminoacyl-tRNA biosynthesis	KEGG
cpic01100	Metabolic pathways	KEGG

Protein-Protein Interaction (PPI)

Clik here to download ENSCPBG00000001138's network

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
ENSCPBG00000001138	EPRS	99	88.155	ENSG00000136628	EPRS	99	82.098	Homo_sapiens
ENSCPBG00000001138	EPRS	98	81.093	ENSAPOG00000002084	eprs	87	81.093	Acanthochromis_polyacanthus
ENSCPBG00000001138	EPRS	99	86.333	ENSAMEG00000009983	EPRS	99	79.401	Ailuropoda_melanoleuca
ENSCPBG00000001138	EPRS	98	80.410	ENSACIG00000007907	eprs	90	80.410	Amphilophus_citrinellus
ENSCPBG00000001138	EPRS	98	77.904	ENSAOCG00000023555	eprs	90	77.904	Amphiprion_ocellaris
ENSCPBG00000001138	EPRS	98	74.715	ENSAPEG00000001630	eprs	99	68.084	Amphiprion_percula
ENSCPBG00000001138	EPRS	98	81.321	ENSATEG00000006493	eprs	100	69.298	Anabas_testudineus
ENSCPBG00000001138	EPRS	99	91.572	ENSAPLG00000003240	EPRS	91	91.572	Anas_platyrhynchos
ENSCPBG00000001138	EPRS	100	91.116	ENSACAG00000001213	EPRS	100	84.128	Anolis_carolinensis
ENSCPBG00000001138	EPRS	99	78.077	ENSANAG00000022208	EPRS	97	84.597	Aotus_nancymaae
ENSCPBG00000001138	EPRS	98	81.093	ENSACLG00000020228	eprs	95	81.093	Astatotilapia_calliptera
ENSCPBG00000001138	EPRS	98	80.638	ENSAMXG00000001984	eprs	90	80.638	Astyanax_mexicanus
ENSCPBG00000001138	EPRS	100	87.472	ENSBTAG00000006898	EPRS	99	83.344	Bos_taurus
ENSCPBG00000001138	EPRS	58	63.524	WBGene00001337	ears-1	98	46.859	Caenorhabditis_elegans
ENSCPBG00000001138	EPRS	99	87.927	ENSCJAG00000003967	EPRS	97	84.123	Callithrix_jacchus
ENSCPBG00000001138	EPRS	99	86.560	ENSCAFG00000010873	EPRS	99	82.917	Canis_familiaris
ENSCPBG00000001138	EPRS	99	86.560	ENSCAFG00020004097	EPRS	99	82.983	Canis_lupus_dingo
ENSCPBG00000001138	EPRS	100	87.244	ENSCHIG00000021232	EPRS	99	83.071	Capra_hircus
ENSCPBG00000001138	EPRS	99	85.877	ENSTSYG00000007633	EPRS	97	84.834	Carlito_syrichta
ENSCPBG00000001138	EPRS	99	83.599	ENSCAPG00000005713	EPRS	94	83.599	Cavia_aperea
ENSCPBG00000001138	EPRS	99	83.599	ENSCPOG00000003586	EPRS	99	78.609	Cavia_porcellus
ENSCPBG00000001138	EPRS	99	88.383	ENSCCAG00000028256	EPRS	100	79.685	Cebus_capucinus
ENSCPBG00000001138	EPRS	97	80.978	ENSCATG00000045050	EPRS	99	81.242	Cercocebus_atys
ENSCPBG00000001138	EPRS	99	85.877	ENSCLAG00000003851	EPRS	99	80.039	Chinchilla_lanigera
ENSCPBG00000001138	EPRS	99	87.927	ENSCSAG00000015776	EPRS	99	82.218	Chlorocebus_sabaeus
ENSCPBG00000001138	EPRS	91	82.143	ENSCHOG00000000140	EPRS	95	82.143	Choloepus_hoffmanni
ENSCPBG00000001138	EPRS	98	69.091	ENSCING00000007544	-	92	59.910	Ciona_intestinalis
ENSCPBG00000001138	EPRS	97	70.975	ENSCSAVG00000007760	-	100	66.102	Ciona_savignyi
ENSCPBG00000001138	EPRS	99	88.383	ENSCANG00000022575	EPRS	99	82.087	Colobus_angolensis_palliatus
ENSCPBG00000001138	EPRS	99	85.194	ENSCGRG00001009885	EPRS	99	81.180	Cricetulus_griseus_chok1gshd
ENSCPBG00000001138	EPRS	99	85.194	ENSCGRG00000010134	Eprs	100	81.337	Cricetulus_griseus_crigri
ENSCPBG00000001138	EPRS	98	80.182	ENSCSEG00000010266	eprs	91	80.182	Cynoglossus_semilaevis
ENSCPBG00000001138	EPRS	98	82.460	ENSCVAG00000005265	eprs	92	82.460	Cyprinodon_variegatus
ENSCPBG00000001138	EPRS	99	82.232	ENSDARG00000060494	eprs	100	69.840	Danio_rerio
ENSCPBG00000001138	EPRS	99	87.699	ENSDNOG00000004016	EPRS	99	82.164	Dasypus_novemcinctus
ENSCPBG00000001138	EPRS	99	86.788	ENSDORG00000004335	Eprs	99	80.482	Dipodomys_ordii
ENSCPBG00000001138	EPRS	99	57.012	FBgn0005674	GluProRS	99	56.937	Drosophila_melanogaster
ENSCPBG00000001138	EPRS	95	94.595	ENSETEG00000019858	EPRS	80	94.595	Echinops_telfairi
ENSCPBG00000001138	EPRS	100	87.016	ENSEASG00005003530	EPRS	99	83.016	Equus_asinus_asinus
ENSCPBG00000001138	EPRS	100	87.016	ENSECAG00000000700	EPRS	99	83.005	Equus_caballus
ENSCPBG00000001138	EPRS	92	85.877	ENSEEUG00000001019	EPRS	87	85.921	Erinaceus_europaeus
ENSCPBG00000001138	EPRS	98	81.321	ENSELUG00000019744	eprs	100	70.033	Esox_lucius
ENSCPBG00000001138	EPRS	99	86.788	ENSFCAG00000002590	EPRS	99	82.283	Felis_catus
ENSCPBG00000001138	EPRS	100	89.294	ENSFALG00000012160	EPRS	95	89.294	Ficedula_albicollis
ENSCPBG00000001138	EPRS	99	83.371	ENSFDAG00000021307	EPRS	100	78.543	Fukomys_damarensis
ENSCPBG00000001138	EPRS	98	81.549	ENSFHEG00000001390	eprs	92	81.549	Fundulus_heteroclitus
ENSCPBG00000001138	EPRS	90	78.360	ENSGMOG00000009984	eprs	93	78.631	Gadus_morhua
ENSCPBG00000001138	EPRS	100	91.116	ENSGALG00000009598	EPRS	99	86.231	Gallus_gallus
ENSCPBG00000001138	EPRS	98	77.449	ENSGAFG00000000716	eprs	100	67.927	Gambusia_affinis
ENSCPBG00000001138	EPRS	98	79.954	ENSGACG00000014823	eprs	93	79.954	Gasterosteus_aculeatus
ENSCPBG00000001138	EPRS	100	98.405	ENSGAGG00000017949	EPRS	100	95.396	Gopherus_agassizii
ENSCPBG00000001138	EPRS	99	88.155	ENSGGOG00000012404	EPRS	99	82.087	Gorilla_gorilla
ENSCPBG00000001138	EPRS	98	81.321	ENSHBUG00000019547	eprs	90	81.321	Haplochromis_burtoni
ENSCPBG00000001138	EPRS	99	84.282	ENSHGLG00000001974	EPRS	99	78.989	Heterocephalus_glaber_female
ENSCPBG00000001138	EPRS	99	84.282	ENSHGLG00100003853	EPRS	99	78.858	Heterocephalus_glaber_male
ENSCPBG00000001138	EPRS	98	80.638	ENSHCOG00000001640	eprs	98	73.962	Hippocampus_comes
ENSCPBG00000001138	EPRS	98	81.777	ENSIPUG00000023784	eprs	90	81.777	Ictalurus_punctatus
ENSCPBG00000001138	EPRS	99	84.510	ENSSTOG00000011870	EPRS	98	81.271	Ictidomys_tridecemlineatus
ENSCPBG00000001138	EPRS	99	84.282	ENSJJAG00000022291	Eprs	99	79.396	Jaculus_jaculus
ENSCPBG00000001138	EPRS	98	81.093	ENSKMAG00000001181	eprs	90	81.093	Kryptolebias_marmoratus
ENSCPBG00000001138	EPRS	98	80.866	ENSLBEG00000016168	eprs	92	80.866	Labrus_bergylta
ENSCPBG00000001138	EPRS	99	85.421	ENSLACG00000011773	EPRS	100	78.006	Latimeria_chalumnae
ENSCPBG00000001138	EPRS	99	87.244	ENSLAFG00000004981	EPRS	99	81.115	Loxodonta_africana
ENSCPBG00000001138	EPRS	99	88.610	ENSMFAG00000041902	EPRS	99	82.218	Macaca_fascicularis
ENSCPBG00000001138	EPRS	99	88.610	ENSMMUG00000008722	EPRS	97	84.597	Macaca_mulatta
ENSCPBG00000001138	EPRS	99	88.610	ENSMNEG00000023887	EPRS	99	82.677	Macaca_nemestrina
ENSCPBG00000001138	EPRS	99	88.155	ENSMLEG00000032585	EPRS	91	88.155	Mandrillus_leucophaeus
ENSCPBG00000001138	EPRS	98	79.954	ENSMAMG00000006099	eprs	100	70.298	Mastacembelus_armatus
ENSCPBG00000001138	EPRS	98	70.843	ENSMZEG00005004950	eprs	99	67.120	Maylandia_zebra
ENSCPBG00000001138	EPRS	100	90.888	ENSMGAG00000009215	EPRS	99	84.897	Meleagris_gallopavo
ENSCPBG00000001138	EPRS	99	84.282	ENSMAUG00000012512	Eprs	99	80.131	Mesocricetus_auratus
ENSCPBG00000001138	EPRS	99	86.333	ENSMICG00000038220	EPRS	99	81.967	Microcebus_murinus
ENSCPBG00000001138	EPRS	99	84.055	ENSMOCG00000012748	Eprs	99	80.787	Microtus_ochrogaster
ENSCPBG00000001138	EPRS	98	81.093	ENSMMOG00000015737	eprs	100	66.950	Mola_mola
ENSCPBG00000001138	EPRS	99	88.155	ENSMODG00000004398	EPRS	99	81.659	Monodelphis_domestica
ENSCPBG00000001138	EPRS	98	80.866	ENSMALG00000014670	eprs	90	80.866	Monopterus_albus
ENSCPBG00000001138	EPRS	99	84.055	MGP_CAROLIEiJ_G0015023	Eprs	99	80.197	Mus_caroli
ENSCPBG00000001138	EPRS	99	84.055	ENSMUSG00000026615	Eprs	99	80.459	Mus_musculus
ENSCPBG00000001138	EPRS	99	84.282	MGP_PahariEiJ_G0028255	Eprs	99	80.131	Mus_pahari
ENSCPBG00000001138	EPRS	99	84.055	MGP_SPRETEiJ_G0015823	Eprs	99	80.262	Mus_spretus
ENSCPBG00000001138	EPRS	99	85.649	ENSMPUG00000004156	EPRS	99	82.589	Mustela_putorius_furo
ENSCPBG00000001138	EPRS	100	86.788	ENSMLUG00000007253	-	93	86.788	Myotis_lucifugus
ENSCPBG00000001138	EPRS	99	85.421	ENSNGAG00000009088	Eprs	99	80.656	Nannospalax_galili
ENSCPBG00000001138	EPRS	98	73.349	ENSNBRG00000006007	eprs	99	67.607	Neolamprologus_brichardi
ENSCPBG00000001138	EPRS	99	87.927	ENSNLEG00000013988	EPRS	99	82.353	Nomascus_leucogenys
ENSCPBG00000001138	EPRS	89	80.463	ENSMEUG00000003870	EPRS	87	76.981	Notamacropus_eugenii
ENSCPBG00000001138	EPRS	89	86.052	ENSOPRG00000001503	EPRS	81	86.052	Ochotona_princeps
ENSCPBG00000001138	EPRS	99	84.510	ENSODEG00000014961	EPRS	99	77.895	Octodon_degus
ENSCPBG00000001138	EPRS	98	81.549	ENSONIG00000003054	eprs	90	81.549	Oreochromis_niloticus
ENSCPBG00000001138	EPRS	99	88.155	ENSOANG00000001424	-	100	81.718	Ornithorhynchus_anatinus
ENSCPBG00000001138	EPRS	99	86.788	ENSOCUG00000012039	EPRS	99	79.387	Oryctolagus_cuniculus
ENSCPBG00000001138	EPRS	98	81.777	ENSORLG00000014819	eprs	100	71.022	Oryzias_latipes
ENSCPBG00000001138	EPRS	98	81.549	ENSORLG00020018395	eprs	100	71.410	Oryzias_latipes_hni
ENSCPBG00000001138	EPRS	98	81.321	ENSORLG00015015996	eprs	100	71.447	Oryzias_latipes_hsok
ENSCPBG00000001138	EPRS	95	81.538	ENSOMEG00000018971	eprs	100	74.960	Oryzias_melastigma
ENSCPBG00000001138	EPRS	99	86.333	ENSOGAG00000005285	EPRS	100	81.684	Otolemur_garnettii
ENSCPBG00000001138	EPRS	100	87.699	ENSOARG00000011829	EPRS	99	83.016	Ovis_aries
ENSCPBG00000001138	EPRS	99	88.383	ENSPPAG00000036757	EPRS	99	82.743	Pan_paniscus
ENSCPBG00000001138	EPRS	99	86.788	ENSPPRG00000012626	EPRS	99	81.955	Panthera_pardus
ENSCPBG00000001138	EPRS	99	86.788	ENSPTIG00000012991	EPRS	99	81.362	Panthera_tigris_altaica
ENSCPBG00000001138	EPRS	99	88.383	ENSPTRG00000001995	EPRS	99	82.349	Pan_troglodytes
ENSCPBG00000001138	EPRS	99	88.155	ENSPANG00000004476	EPRS	99	81.800	Papio_anubis
ENSCPBG00000001138	EPRS	98	81.093	ENSPKIG00000023015	eprs	93	81.093	Paramormyrops_kingsleyae
ENSCPBG00000001138	EPRS	100	94.989	ENSPSIG00000010709	EPRS	100	91.771	Pelodiscus_sinensis
ENSCPBG00000001138	EPRS	98	74.715	ENSPMGG00000023169	eprs	99	67.782	Periophthalmus_magnuspinnatus
ENSCPBG00000001138	EPRS	99	79.041	ENSPEMG00000017287	Eprs	99	79.041	Peromyscus_maniculatus_bairdii
ENSCPBG00000001138	EPRS	99	86.333	ENSPCIG00000009736	EPRS	99	81.639	Phascolarctos_cinereus
ENSCPBG00000001138	EPRS	99	82.005	ENSPFOG00000017972	eprs	100	71.728	Poecilia_formosa
ENSCPBG00000001138	EPRS	65	68.070	ENSPLAG00000016198	EPRS	99	68.458	Poecilia_latipinna
ENSCPBG00000001138	EPRS	67	82.005	ENSPMEG00000022964	eprs	100	72.585	Poecilia_mexicana
ENSCPBG00000001138	EPRS	65	68.028	ENSPMEG00000001890	EPRS	99	68.998	Poecilia_mexicana
ENSCPBG00000001138	EPRS	98	81.549	ENSPREG00000004302	eprs	89	81.549	Poecilia_reticulata
ENSCPBG00000001138	EPRS	99	81.321	ENSPPYG00000000210	EPRS	99	81.549	Pongo_abelii
ENSCPBG00000001138	EPRS	85	83.945	ENSPCAG00000015737	EPRS	88	82.278	Procavia_capensis
ENSCPBG00000001138	EPRS	99	80.182	ENSPCOG00000004103	EPRS	97	81.406	Propithecus_coquereli
ENSCPBG00000001138	EPRS	99	84.738	ENSPVAG00000013147	EPRS	99	80.262	Pteropus_vampyrus
ENSCPBG00000001138	EPRS	98	81.549	ENSPNYG00000007931	eprs	90	81.549	Pundamilia_nyererei
ENSCPBG00000001138	EPRS	99	80.638	ENSPNAG00000003344	eprs	94	80.638	Pygocentrus_nattereri
ENSCPBG00000001138	EPRS	99	83.599	ENSRNOG00000002393	Eprs	99	79.279	Rattus_norvegicus
ENSCPBG00000001138	EPRS	99	88.610	ENSRBIG00000034767	EPRS	100	83.121	Rhinopithecus_bieti
ENSCPBG00000001138	EPRS	99	88.610	ENSRROG00000039885	EPRS	99	82.283	Rhinopithecus_roxellana
ENSCPBG00000001138	EPRS	99	88.383	ENSSBOG00000029926	EPRS	99	82.612	Saimiri_boliviensis_boliviensis
ENSCPBG00000001138	EPRS	89	88.435	ENSSHAG00000002391	EPRS	99	82.417	Sarcophilus_harrisii
ENSCPBG00000001138	EPRS	99	82.005	ENSSFOG00015007233	eprs	100	70.649	Scleropages_formosus
ENSCPBG00000001138	EPRS	98	80.410	ENSSMAG00000011150	eprs	100	69.446	Scophthalmus_maximus
ENSCPBG00000001138	EPRS	98	81.321	ENSSDUG00000008910	eprs	91	81.321	Seriola_dumerili
ENSCPBG00000001138	EPRS	98	81.321	ENSSLDG00000015153	eprs	100	73.362	Seriola_lalandi_dorsalis
ENSCPBG00000001138	EPRS	95	76.823	ENSSARG00000012244	EPRS	99	76.891	Sorex_araneus
ENSCPBG00000001138	EPRS	100	83.827	ENSSPUG00000009384	EPRS	97	81.057	Sphenodon_punctatus
ENSCPBG00000001138	EPRS	98	81.321	ENSSPAG00000010408	eprs	100	71.745	Stegastes_partitus
ENSCPBG00000001138	EPRS	99	87.472	ENSSSCG00000010821	EPRS	99	83.136	Sus_scrofa
ENSCPBG00000001138	EPRS	100	85.877	ENSTGUG00000002796	EPRS	100	84.712	Taeniopygia_guttata
ENSCPBG00000001138	EPRS	98	80.410	ENSTRUG00000006901	eprs	91	80.410	Takifugu_rubripes
ENSCPBG00000001138	EPRS	65	65.670	ENSTNIG00000011902	eprs	99	65.670	Tetraodon_nigroviridis
ENSCPBG00000001138	EPRS	99	86.788	ENSTBEG00000008365	EPRS	99	82.230	Tupaia_belangeri
ENSCPBG00000001138	EPRS	99	85.877	ENSTTRG00000013013	EPRS	99	82.283	Tursiops_truncatus
ENSCPBG00000001138	EPRS	99	86.105	ENSUAMG00000020817	EPRS	99	82.457	Ursus_americanus
ENSCPBG00000001138	EPRS	99	86.105	ENSUMAG00000020899	EPRS	100	82.720	Ursus_maritimus
ENSCPBG00000001138	EPRS	99	76.806	ENSVPAG00000008903	EPRS	99	72.586	Vicugna_pacos
ENSCPBG00000001138	EPRS	99	85.877	ENSVVUG00000012841	EPRS	77	82.700	Vulpes_vulpes
ENSCPBG00000001138	EPRS	99	87.273	ENSXETG00000002724	eprs	91	87.273	Xenopus_tropicalis
ENSCPBG00000001138	EPRS	91	81.777	ENSXCOG00000018038	eprs	92	81.777	Xiphophorus_couchianus
ENSCPBG00000001138	EPRS	98	81.777	ENSXMAG00000011690	eprs	91	81.777	Xiphophorus_maculatus