EuRBPDB

Description

RBDs

Transcripts

Gene Model

PPI

Paralogs

Orthologs

Description

Ensembl ID: ENSEBUG00000008179 (Gene tree)
Gene ID: -
Gene Symbol: -
Alias: zgc:91918
Full Name: -
Gene Type: protein_coding
Species: Eptatretus_burgeri
Status: putative
Strand: Minus strand
Length: 335763 bases
Position: chrFYBX02009559.1:1636340-1972102
Accession: -
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
ENSEBUP00000012949	RRM_5	PF13893.6	2.5e-12	1	2
ENSEBUP00000012949	RRM_5	PF13893.6	2.5e-12	2	2
ENSEBUP00000012896	RRM_5	PF13893.6	3e-12	1	2
ENSEBUP00000012896	RRM_5	PF13893.6	3e-12	2	2
ENSEBUP00000012920	RRM_5	PF13893.6	3e-12	1	2
ENSEBUP00000012920	RRM_5	PF13893.6	3e-12	2	2
ENSEBUP00000012931	RRM_5	PF13893.6	3e-12	1	2
ENSEBUP00000012931	RRM_5	PF13893.6	3e-12	2	2
ENSEBUP00000012909	RRM_5	PF13893.6	3.3e-12	1	2
ENSEBUP00000012909	RRM_5	PF13893.6	3.3e-12	2	2
ENSEBUP00000012949	RRM_1	PF00076.22	2.3e-64	1	3
ENSEBUP00000012949	RRM_1	PF00076.22	2.3e-64	2	3
ENSEBUP00000012949	RRM_1	PF00076.22	2.3e-64	3	3
ENSEBUP00000012896	RRM_1	PF00076.22	2.7e-64	1	3
ENSEBUP00000012896	RRM_1	PF00076.22	2.7e-64	2	3
ENSEBUP00000012896	RRM_1	PF00076.22	2.7e-64	3	3
ENSEBUP00000012920	RRM_1	PF00076.22	2.7e-64	1	3
ENSEBUP00000012920	RRM_1	PF00076.22	2.7e-64	2	3
ENSEBUP00000012920	RRM_1	PF00076.22	2.7e-64	3	3
ENSEBUP00000012931	RRM_1	PF00076.22	2.7e-64	1	3
ENSEBUP00000012931	RRM_1	PF00076.22	2.7e-64	2	3
ENSEBUP00000012931	RRM_1	PF00076.22	2.7e-64	3	3
ENSEBUP00000012909	RRM_1	PF00076.22	3e-64	1	3
ENSEBUP00000012909	RRM_1	PF00076.22	3e-64	2	3
ENSEBUP00000012909	RRM_1	PF00076.22	3e-64	3	3

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
ENSEBUT00000013485	-	1620	-	ENSEBUP00000012909	377 (aa)	-	-
ENSEBUT00000013507	-	2198	-	ENSEBUP00000012931	369 (aa)	-	-
ENSEBUT00000013525	-	2159	-	ENSEBUP00000012949	356 (aa)	-	-
ENSEBUT00000013496	-	1455	-	ENSEBUP00000012920	369 (aa)	-	-
ENSEBUT00000013473	-	1450	-	ENSEBUP00000012896	369 (aa)	-	-

Gene Model

Click here to download ENSEBUG00000008179's gene model file

Protein-Protein Interaction (PPI)

Clik here to download ENSEBUG00000008179's network

Paralogs

Ensembl ID	Gene Symbol	Coverage	Identiy	Paralog	Gene Symbol	Coverage	Identiy
ENSEBUG00000008179	-	71	41.463	ENSEBUG00000002471	-	53	41.463
ENSEBUG00000008179	-	89	83.860	ENSEBUG00000016008	elavl2	97	83.860
ENSEBUG00000008179	-	73	40.964	ENSEBUG00000007835	-	63	40.964
ENSEBUG00000008179	-	69	39.744	ENSEBUG00000001843	tia1l	55	39.744
ENSEBUG00000008179	-	76	41.667	ENSEBUG00000008128	-	53	41.667
ENSEBUG00000008179	-	85	30.128	ENSEBUG00000007042	rbm45	90	30.128
ENSEBUG00000008179	-	87	37.349	ENSEBUG00000000992	pabpc4	62	37.349
ENSEBUG00000008179	-	57	35.096	ENSEBUG00000003357	-	50	37.017
ENSEBUG00000008179	-	87	33.333	ENSEBUG00000013782	pabpc4	62	31.126
ENSEBUG00000008179	-	86	67.638	ENSEBUG00000000352	-	95	67.638
ENSEBUG00000008179	-	74	42.683	ENSEBUG00000011955	CELF5	54	42.683

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
ENSEBUG00000008179	-	100	84.011	ENSG00000107105	ELAVL2	100	91.111	Homo_sapiens
ENSEBUG00000008179	-	100	80.541	ENSAPOG00000022121	ELAVL2	94	80.267	Acanthochromis_polyacanthus
ENSEBUG00000008179	-	100	83.784	ENSAMEG00000007555	ELAVL2	94	82.667	Ailuropoda_melanoleuca
ENSEBUG00000008179	-	100	83.243	ENSACIG00000010855	ELAVL2	99	90.476	Amphilophus_citrinellus
ENSEBUG00000008179	-	100	83.514	ENSAOCG00000019468	ELAVL2	94	83.200	Amphiprion_ocellaris
ENSEBUG00000008179	-	100	83.514	ENSAPEG00000023474	ELAVL2	94	83.200	Amphiprion_percula
ENSEBUG00000008179	-	100	84.011	ENSATEG00000018073	ELAVL2	100	84.011	Anabas_testudineus
ENSEBUG00000008179	-	100	83.243	ENSAPLG00000010429	ELAVL2	94	81.867	Anas_platyrhynchos
ENSEBUG00000008179	-	100	84.054	ENSACAG00000017521	ELAVL2	88	82.933	Anolis_carolinensis
ENSEBUG00000008179	-	100	84.011	ENSANAG00000032553	ELAVL2	99	82.447	Aotus_nancymaae
ENSEBUG00000008179	-	100	83.243	ENSACLG00000002715	ELAVL2	94	82.933	Astatotilapia_calliptera
ENSEBUG00000008179	-	100	84.595	ENSAMXG00000036990	ELAVL2	94	83.200	Astyanax_mexicanus
ENSEBUG00000008179	-	100	83.784	ENSBTAG00000046348	ELAVL2	100	82.973	Bos_taurus
ENSEBUG00000008179	-	100	83.784	ENSCJAG00000008064	ELAVL2	99	82.973	Callithrix_jacchus
ENSEBUG00000008179	-	100	84.011	ENSCAFG00000001688	ELAVL2	94	82.888	Canis_familiaris
ENSEBUG00000008179	-	100	84.011	ENSCAFG00020013836	ELAVL2	92	90.826	Canis_lupus_dingo
ENSEBUG00000008179	-	100	84.011	ENSCHIG00000025894	ELAVL2	100	83.198	Capra_hircus
ENSEBUG00000008179	-	100	83.784	ENSTSYG00000013066	ELAVL2	94	82.667	Carlito_syrichta
ENSEBUG00000008179	-	90	85.304	ENSCAPG00000003609	ELAVL2	100	85.304	Cavia_aperea
ENSEBUG00000008179	-	100	83.740	ENSCPOG00000010472	ELAVL2	98	81.963	Cavia_porcellus
ENSEBUG00000008179	-	100	83.784	ENSCCAG00000024508	ELAVL2	98	82.667	Cebus_capucinus
ENSEBUG00000008179	-	100	84.011	ENSCATG00000012522	ELAVL2	98	82.011	Cercocebus_atys
ENSEBUG00000008179	-	100	83.784	ENSCLAG00000003388	ELAVL2	98	82.011	Chinchilla_lanigera
ENSEBUG00000008179	-	100	84.011	ENSCSAG00000009718	ELAVL2	94	82.888	Chlorocebus_sabaeus
ENSEBUG00000008179	-	100	85.470	ENSCHOG00000000635	ELAVL2	75	90.083	Choloepus_hoffmanni
ENSEBUG00000008179	-	100	84.054	ENSCPBG00000007841	ELAVL2	100	83.243	Chrysemys_picta_bellii
ENSEBUG00000008179	-	87	62.712	ENSCING00000004712	-	92	62.745	Ciona_intestinalis
ENSEBUG00000008179	-	85	70.000	ENSCSAVG00000003440	-	99	68.371	Ciona_savignyi
ENSEBUG00000008179	-	100	84.011	ENSCANG00000043179	ELAVL2	100	83.198	Colobus_angolensis_palliatus
ENSEBUG00000008179	-	100	84.054	ENSCGRG00001024409	Elavl2	100	82.584	Cricetulus_griseus_chok1gshd
ENSEBUG00000008179	-	100	84.054	ENSCGRG00000000487	Elavl2	100	83.198	Cricetulus_griseus_crigri
ENSEBUG00000008179	-	100	83.784	ENSCSEG00000006335	elavl2	94	83.467	Cynoglossus_semilaevis
ENSEBUG00000008179	-	100	83.243	ENSCVAG00000013913	ELAVL2	94	82.667	Cyprinodon_variegatus
ENSEBUG00000008179	-	100	82.656	ENSDARG00000040732	elavl2	100	86.782	Danio_rerio
ENSEBUG00000008179	-	100	83.784	ENSDNOG00000035657	ELAVL2	94	82.667	Dasypus_novemcinctus
ENSEBUG00000008179	-	100	84.011	ENSDORG00000010437	Elavl2	98	82.228	Dipodomys_ordii
ENSEBUG00000008179	-	87	62.985	FBgn0010263	Rbp9	76	63.284	Drosophila_melanogaster
ENSEBUG00000008179	-	88	64.412	FBgn0086675	fne	94	64.095	Drosophila_melanogaster
ENSEBUG00000008179	-	100	84.011	ENSETEG00000006134	ELAVL2	94	82.933	Echinops_telfairi
ENSEBUG00000008179	-	100	84.011	ENSEASG00005004697	ELAVL2	94	82.888	Equus_asinus_asinus
ENSEBUG00000008179	-	100	84.011	ENSECAG00000020142	ELAVL2	95	82.584	Equus_caballus
ENSEBUG00000008179	-	100	69.648	ENSEEUG00000008733	ELAVL2	95	68.533	Erinaceus_europaeus
ENSEBUG00000008179	-	100	84.595	ENSELUG00000010026	ELAVL2	94	84.267	Esox_lucius
ENSEBUG00000008179	-	100	84.011	ENSFCAG00000011976	ELAVL2	100	82.584	Felis_catus
ENSEBUG00000008179	-	100	82.973	ENSFALG00000007363	ELAVL2	94	81.867	Ficedula_albicollis
ENSEBUG00000008179	-	100	84.282	ENSFDAG00000018820	ELAVL2	98	82.493	Fukomys_damarensis
ENSEBUG00000008179	-	100	80.541	ENSFHEG00000010307	ELAVL2	94	80.000	Fundulus_heteroclitus
ENSEBUG00000008179	-	100	83.514	ENSGMOG00000011182	ELAVL2	100	83.514	Gadus_morhua
ENSEBUG00000008179	-	100	82.703	ENSGALG00000015799	ELAVL2	94	80.789	Gallus_gallus
ENSEBUG00000008179	-	100	83.784	ENSGAFG00000016517	ELAVL2	86	83.200	Gambusia_affinis
ENSEBUG00000008179	-	100	83.514	ENSGACG00000019484	elavl2	100	83.514	Gasterosteus_aculeatus
ENSEBUG00000008179	-	100	84.054	ENSGAGG00000022382	ELAVL2	100	83.243	Gopherus_agassizii
ENSEBUG00000008179	-	51	85.965	ENSGAGG00000005858	-	99	85.965	Gopherus_agassizii
ENSEBUG00000008179	-	100	83.784	ENSGGOG00000003678	ELAVL2	100	82.353	Gorilla_gorilla
ENSEBUG00000008179	-	100	83.243	ENSHBUG00000005968	ELAVL2	94	82.933	Haplochromis_burtoni
ENSEBUG00000008179	-	100	84.011	ENSHGLG00000005423	ELAVL2	100	83.198	Heterocephalus_glaber_female
ENSEBUG00000008179	-	100	84.011	ENSHGLG00100002344	ELAVL2	100	83.198	Heterocephalus_glaber_male
ENSEBUG00000008179	-	100	77.297	ENSHCOG00000005961	elavl2	94	77.067	Hippocampus_comes
ENSEBUG00000008179	-	98	83.657	ENSIPUG00000012337	elavl2	88	83.060	Ictalurus_punctatus
ENSEBUG00000008179	-	100	84.282	ENSSTOG00000007792	ELAVL2	98	83.155	Ictidomys_tridecemlineatus
ENSEBUG00000008179	-	100	84.011	ENSJJAG00000008247	Elavl2	98	82.228	Jaculus_jaculus
ENSEBUG00000008179	-	100	83.243	ENSKMAG00000006299	ELAVL2	94	82.667	Kryptolebias_marmoratus
ENSEBUG00000008179	-	100	83.514	ENSLBEG00000027924	ELAVL2	96	82.913	Labrus_bergylta
ENSEBUG00000008179	-	100	84.011	ENSLACG00000010888	ELAVL2	93	89.062	Latimeria_chalumnae
ENSEBUG00000008179	-	100	83.784	ENSLOCG00000012575	ELAVL2	94	83.467	Lepisosteus_oculatus
ENSEBUG00000008179	-	100	83.784	ENSLAFG00000009938	ELAVL2	98	82.493	Loxodonta_africana
ENSEBUG00000008179	-	100	84.011	ENSMFAG00000039694	ELAVL2	100	82.973	Macaca_fascicularis
ENSEBUG00000008179	-	100	84.011	ENSMMUG00000010038	ELAVL2	99	82.275	Macaca_mulatta
ENSEBUG00000008179	-	100	84.011	ENSMNEG00000032526	ELAVL2	100	82.973	Macaca_nemestrina
ENSEBUG00000008179	-	51	90.435	ENSMLEG00000027781	-	100	90.435	Mandrillus_leucophaeus
ENSEBUG00000008179	-	100	83.198	ENSMAMG00000004806	ELAVL2	100	83.198	Mastacembelus_armatus
ENSEBUG00000008179	-	100	83.243	ENSMZEG00005000778	ELAVL2	94	82.933	Maylandia_zebra
ENSEBUG00000008179	-	85	61.702	ENSMGAG00000008665	ELAVL2	100	61.702	Meleagris_gallopavo
ENSEBUG00000008179	-	100	84.011	ENSMAUG00000011955	Elavl2	100	82.353	Mesocricetus_auratus
ENSEBUG00000008179	-	100	84.011	ENSMICG00000001558	ELAVL2	96	82.493	Microcebus_murinus
ENSEBUG00000008179	-	100	84.011	ENSMOCG00000016960	Elavl2	98	82.228	Microtus_ochrogaster
ENSEBUG00000008179	-	100	83.243	ENSMMOG00000012031	ELAVL2	100	83.243	Mola_mola
ENSEBUG00000008179	-	100	84.054	ENSMODG00000003936	ELAVL2	94	82.933	Monodelphis_domestica
ENSEBUG00000008179	-	100	83.469	ENSMALG00000003411	elavl2	90	83.155	Monopterus_albus
ENSEBUG00000008179	-	100	84.011	MGP_CAROLIEiJ_G0026159	Elavl2	100	82.973	Mus_caroli
ENSEBUG00000008179	-	100	84.011	ENSMUSG00000008489	Elavl2	100	83.198	Mus_musculus
ENSEBUG00000008179	-	100	84.011	MGP_PahariEiJ_G0028496	Elavl2	100	82.973	Mus_pahari
ENSEBUG00000008179	-	100	84.011	MGP_SPRETEiJ_G0027124	Elavl2	100	82.973	Mus_spretus
ENSEBUG00000008179	-	100	84.011	ENSMPUG00000015967	ELAVL2	94	82.888	Mustela_putorius_furo
ENSEBUG00000008179	-	100	80.940	ENSMLUG00000014210	ELAVL2	99	79.284	Myotis_lucifugus
ENSEBUG00000008179	-	100	83.740	ENSNGAG00000014535	Elavl2	98	81.963	Nannospalax_galili
ENSEBUG00000008179	-	100	74.373	ENSNBRG00000010284	ELAVL2	100	74.373	Neolamprologus_brichardi
ENSEBUG00000008179	-	90	91.815	ENSNBRG00000023764	-	100	90.521	Neolamprologus_brichardi
ENSEBUG00000008179	-	100	84.011	ENSNLEG00000016382	ELAVL2	100	83.198	Nomascus_leucogenys
ENSEBUG00000008179	-	100	75.068	ENSMEUG00000006499	ELAVL2	94	73.867	Notamacropus_eugenii
ENSEBUG00000008179	-	100	84.011	ENSOPRG00000012874	ELAVL2	94	82.667	Ochotona_princeps
ENSEBUG00000008179	-	100	84.011	ENSODEG00000002531	ELAVL2	100	83.198	Octodon_degus
ENSEBUG00000008179	-	100	81.698	ENSONIG00000007552	ELAVL2	100	81.414	Oreochromis_niloticus
ENSEBUG00000008179	-	100	78.706	ENSOANG00000009634	ELAVL2	91	77.660	Ornithorhynchus_anatinus
ENSEBUG00000008179	-	100	84.011	ENSOCUG00000005891	ELAVL2	94	82.888	Oryctolagus_cuniculus
ENSEBUG00000008179	-	100	83.514	ENSORLG00000004217	ELAVL2	100	81.894	Oryzias_latipes
ENSEBUG00000008179	-	100	83.514	ENSORLG00020016040	ELAVL2	94	83.200	Oryzias_latipes_hni
ENSEBUG00000008179	-	100	83.514	ENSORLG00015013661	ELAVL2	94	83.200	Oryzias_latipes_hsok
ENSEBUG00000008179	-	100	83.514	ENSOMEG00000005178	ELAVL2	94	83.200	Oryzias_melastigma
ENSEBUG00000008179	-	98	80.615	ENSOGAG00000016228	ELAVL2	98	78.916	Otolemur_garnettii
ENSEBUG00000008179	-	100	84.011	ENSOARG00000014465	ELAVL2	98	82.228	Ovis_aries
ENSEBUG00000008179	-	100	84.011	ENSPPAG00000041539	ELAVL2	99	82.713	Pan_paniscus
ENSEBUG00000008179	-	100	83.784	ENSPPRG00000018510	ELAVL2	100	82.584	Panthera_pardus
ENSEBUG00000008179	-	100	84.011	ENSPTIG00000005627	ELAVL2	100	83.198	Panthera_tigris_altaica
ENSEBUG00000008179	-	100	84.011	ENSPTRG00000020830	ELAVL2	100	82.973	Pan_troglodytes
ENSEBUG00000008179	-	100	84.011	ENSPANG00000025263	ELAVL2	98	82.011	Papio_anubis
ENSEBUG00000008179	-	100	82.703	ENSPKIG00000001903	-	100	82.703	Paramormyrops_kingsleyae
ENSEBUG00000008179	-	100	84.054	ENSPKIG00000006187	ELAVL2	96	83.733	Paramormyrops_kingsleyae
ENSEBUG00000008179	-	90	91.318	ENSPSIG00000015330	ELAVL2	100	91.318	Pelodiscus_sinensis
ENSEBUG00000008179	-	100	82.973	ENSPMGG00000022018	elavl2	100	82.973	Periophthalmus_magnuspinnatus
ENSEBUG00000008179	-	100	84.011	ENSPEMG00000014101	Elavl2	98	82.228	Peromyscus_maniculatus_bairdii
ENSEBUG00000008179	-	100	83.784	ENSPCIG00000011696	ELAVL2	98	82.667	Phascolarctos_cinereus
ENSEBUG00000008179	-	100	83.514	ENSPFOG00000003436	ELAVL2	86	82.933	Poecilia_formosa
ENSEBUG00000008179	-	100	83.514	ENSPLAG00000005203	ELAVL2	100	83.243	Poecilia_latipinna
ENSEBUG00000008179	-	100	83.514	ENSPMEG00000021929	ELAVL2	92	82.913	Poecilia_mexicana
ENSEBUG00000008179	-	100	83.514	ENSPREG00000017160	ELAVL2	94	82.933	Poecilia_reticulata
ENSEBUG00000008179	-	100	84.011	ENSPPYG00000019166	ELAVL2	94	82.888	Pongo_abelii
ENSEBUG00000008179	-	100	80.488	ENSPCAG00000014998	ELAVL2	94	79.200	Procavia_capensis
ENSEBUG00000008179	-	100	84.011	ENSPCOG00000022819	ELAVL2	100	82.888	Propithecus_coquereli
ENSEBUG00000008179	-	100	84.011	ENSPVAG00000002688	ELAVL2	94	82.667	Pteropus_vampyrus
ENSEBUG00000008179	-	100	83.243	ENSPNYG00000001514	ELAVL2	99	90.764	Pundamilia_nyererei
ENSEBUG00000008179	-	100	84.054	ENSPNAG00000029068	ELAVL2	94	82.933	Pygocentrus_nattereri
ENSEBUG00000008179	-	100	84.011	ENSRNOG00000006853	Elavl2	100	82.353	Rattus_norvegicus
ENSEBUG00000008179	-	100	84.011	ENSRBIG00000029495	ELAVL2	100	82.584	Rhinopithecus_bieti
ENSEBUG00000008179	-	100	83.784	ENSRROG00000041782	ELAVL2	100	82.973	Rhinopithecus_roxellana
ENSEBUG00000008179	-	100	84.011	ENSSBOG00000028114	ELAVL2	98	82.667	Saimiri_boliviensis_boliviensis
ENSEBUG00000008179	-	100	83.193	ENSSHAG00000006756	ELAVL2	100	84.516	Sarcophilus_harrisii
ENSEBUG00000008179	-	100	80.811	ENSSFOG00015001257	ELAVL2	99	79.101	Scleropages_formosus
ENSEBUG00000008179	-	100	83.784	ENSSFOG00015018252	elavl2	94	83.467	Scleropages_formosus
ENSEBUG00000008179	-	100	83.514	ENSSMAG00000010745	ELAVL2	94	83.200	Scophthalmus_maximus
ENSEBUG00000008179	-	100	83.514	ENSSDUG00000008140	ELAVL2	94	83.200	Seriola_dumerili
ENSEBUG00000008179	-	100	83.514	ENSSLDG00000008455	ELAVL2	100	83.514	Seriola_lalandi_dorsalis
ENSEBUG00000008179	-	100	84.011	ENSSARG00000005702	ELAVL2	94	82.667	Sorex_araneus
ENSEBUG00000008179	-	100	84.054	ENSSPUG00000001246	ELAVL2	100	83.243	Sphenodon_punctatus
ENSEBUG00000008179	-	100	83.514	ENSSPAG00000002388	ELAVL2	100	83.514	Stegastes_partitus
ENSEBUG00000008179	-	100	84.011	ENSSSCG00000005130	ELAVL2	100	83.198	Sus_scrofa
ENSEBUG00000008179	-	99	76.762	ENSTGUG00000001349	ELAVL2	99	76.104	Taeniopygia_guttata
ENSEBUG00000008179	-	100	84.324	ENSTNIG00000011312	elavl2	97	83.155	Tetraodon_nigroviridis
ENSEBUG00000008179	-	100	72.087	ENSTBEG00000017759	ELAVL2	94	70.933	Tupaia_belangeri
ENSEBUG00000008179	-	100	83.740	ENSTTRG00000011656	ELAVL2	94	82.400	Tursiops_truncatus
ENSEBUG00000008179	-	100	83.784	ENSUAMG00000018679	ELAVL2	100	82.540	Ursus_americanus
ENSEBUG00000008179	-	100	84.011	ENSUMAG00000005644	ELAVL2	92	90.826	Ursus_maritimus
ENSEBUG00000008179	-	100	84.011	ENSVPAG00000005109	ELAVL2	94	82.667	Vicugna_pacos
ENSEBUG00000008179	-	100	84.011	ENSVVUG00000021291	ELAVL2	100	82.584	Vulpes_vulpes
ENSEBUG00000008179	-	100	81.501	ENSXETG00000009387	elavl2	100	89.262	Xenopus_tropicalis
ENSEBUG00000008179	-	100	83.193	ENSXCOG00000002464	ELAVL2	99	83.193	Xiphophorus_couchianus
ENSEBUG00000008179	-	100	83.784	ENSXMAG00000029211	ELAVL2	86	83.200	Xiphophorus_maculatus