EuRBPDB

Description

RBDs

Transcripts

Gene Model

PPI

Paralogs

Orthologs

Description

Ensembl ID: ENSETEG00000005732 (Gene tree)
Gene ID: -
Gene Symbol: -
Alias: -
Full Name: ribosomal protein L13a
Gene Type: protein_coding
Species: Echinops_telfairi
Status: putative
Strand: Plus strand
Length: 612 bases
Position: chrscaffold_136149:255-866
Accession: 101650566
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
ENSETEP00000004659	Ribosomal_L13	PF00572.18	2.4e-07	1	1

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
ENSETET00000005730	-	612	XM_004710436	ENSETEP00000004659	203 (aa)	XP_004710493	UPI0000E40DFB

Gene Model

Click here to download ENSETEG00000005732's gene model file

Protein-Protein Interaction (PPI)

Clik here to download ENSETEG00000005732's network

Paralogs

Ensembl ID	Gene Symbol	Coverage	Identiy	Paralog	Gene Symbol	Coverage	Identiy
ENSETEG00000005732	-	100	78.818	ENSETEG00000017076	-	100	78.818

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
ENSETEG00000005732	-	100	94.581	ENSG00000142541	RPL13A	100	94.581	Homo_sapiens
ENSETEG00000005732	-	98	84.925	ENSAPOG00000014084	rpl13a	97	84.925	Acanthochromis_polyacanthus
ENSETEG00000005732	-	100	95.074	ENSAMEG00000014027	RPL13A	100	95.074	Ailuropoda_melanoleuca
ENSETEG00000005732	-	98	84.343	ENSACIG00000004871	rpl13a	99	84.343	Amphilophus_citrinellus
ENSETEG00000005732	-	98	84.925	ENSAOCG00000002123	rpl13a	97	84.925	Amphiprion_ocellaris
ENSETEG00000005732	-	98	84.925	ENSAPEG00000018777	rpl13a	97	84.925	Amphiprion_percula
ENSETEG00000005732	-	98	85.930	ENSATEG00000012828	rpl13a	97	85.930	Anabas_testudineus
ENSETEG00000005732	-	100	87.192	ENSACAG00000017279	RPL13A	100	87.192	Anolis_carolinensis
ENSETEG00000005732	-	100	83.824	ENSANAG00000020868	-	100	83.824	Aotus_nancymaae
ENSETEG00000005732	-	99	81.592	ENSANAG00000021423	-	99	81.592	Aotus_nancymaae
ENSETEG00000005732	-	70	78.378	ENSANAG00000037443	-	100	78.378	Aotus_nancymaae
ENSETEG00000005732	-	70	92.958	ENSANAG00000013769	-	100	92.958	Aotus_nancymaae
ENSETEG00000005732	-	100	94.089	ENSANAG00000030530	RPL13A	100	94.089	Aotus_nancymaae
ENSETEG00000005732	-	98	84.925	ENSACLG00000016552	rpl13a	97	84.925	Astatotilapia_calliptera
ENSETEG00000005732	-	98	84.422	ENSAMXG00000033532	rpl13a	97	84.422	Astyanax_mexicanus
ENSETEG00000005732	-	100	93.596	ENSBTAG00000005296	RPL13A	100	93.596	Bos_taurus
ENSETEG00000005732	-	98	50.754	WBGene00004428	rpl-16	98	50.754	Caenorhabditis_elegans
ENSETEG00000005732	-	100	92.611	ENSCJAG00000040678	-	100	92.611	Callithrix_jacchus
ENSETEG00000005732	-	100	88.177	ENSCJAG00000046649	-	100	88.177	Callithrix_jacchus
ENSETEG00000005732	-	100	95.074	ENSCJAG00000000072	-	100	95.074	Callithrix_jacchus
ENSETEG00000005732	-	82	77.907	ENSCJAG00000008837	-	98	77.907	Callithrix_jacchus
ENSETEG00000005732	-	99	60.500	ENSCAFG00000003946	-	97	60.500	Canis_familiaris
ENSETEG00000005732	-	100	95.074	ENSCAFG00000029892	RPL13A	100	95.074	Canis_familiaris
ENSETEG00000005732	-	100	95.074	ENSCAFG00020024917	-	100	95.074	Canis_lupus_dingo
ENSETEG00000005732	-	100	93.596	ENSCHIG00000021732	-	100	93.596	Capra_hircus
ENSETEG00000005732	-	100	90.640	ENSCHIG00000012581	-	100	90.640	Capra_hircus
ENSETEG00000005732	-	100	94.581	ENSTSYG00000008791	-	100	94.581	Carlito_syrichta
ENSETEG00000005732	-	98	72.362	ENSTSYG00000028115	-	96	74.372	Carlito_syrichta
ENSETEG00000005732	-	98	93.467	ENSCAPG00000007267	RPL13A	100	93.467	Cavia_aperea
ENSETEG00000005732	-	62	95.200	ENSCPOG00000023083	-	59	91.729	Cavia_porcellus
ENSETEG00000005732	-	100	93.596	ENSCPOG00000012417	RPL13A	100	93.596	Cavia_porcellus
ENSETEG00000005732	-	68	88.489	ENSCCAG00000020555	-	100	88.489	Cebus_capucinus
ENSETEG00000005732	-	68	93.525	ENSCCAG00000029220	-	96	93.525	Cebus_capucinus
ENSETEG00000005732	-	100	94.581	ENSCCAG00000022898	RPL13A	100	94.581	Cebus_capucinus
ENSETEG00000005732	-	94	91.623	ENSCCAG00000034479	-	95	91.623	Cebus_capucinus
ENSETEG00000005732	-	100	94.089	ENSCATG00000011576	RPL13A	100	94.089	Cercocebus_atys
ENSETEG00000005732	-	100	93.596	ENSCLAG00000012323	RPL13A	100	93.596	Chinchilla_lanigera
ENSETEG00000005732	-	100	92.611	ENSCSAG00000019054	-	100	92.611	Chlorocebus_sabaeus
ENSETEG00000005732	-	100	85.222	ENSCPBG00000014922	RPL13A	95	85.222	Chrysemys_picta_bellii
ENSETEG00000005732	-	99	60.396	ENSCING00000014892	-	98	60.396	Ciona_intestinalis
ENSETEG00000005732	-	99	62.069	ENSCSAVG00000000666	-	99	62.069	Ciona_savignyi
ENSETEG00000005732	-	100	94.089	ENSCANG00000040885	-	100	94.089	Colobus_angolensis_palliatus
ENSETEG00000005732	-	81	81.325	ENSCANG00000018732	-	100	81.325	Colobus_angolensis_palliatus
ENSETEG00000005732	-	100	93.596	ENSCANG00000000304	-	100	93.596	Colobus_angolensis_palliatus
ENSETEG00000005732	-	100	92.118	ENSCGRG00001018651	-	100	92.118	Cricetulus_griseus_chok1gshd
ENSETEG00000005732	-	100	92.118	ENSCGRG00000005890	-	100	92.118	Cricetulus_griseus_crigri
ENSETEG00000005732	-	99	76.119	ENSCSEG00000002640	rpl13a	98	76.119	Cynoglossus_semilaevis
ENSETEG00000005732	-	99	73.500	ENSCVAG00000011043	rpl13a	97	73.500	Cyprinodon_variegatus
ENSETEG00000005732	-	98	84.422	ENSDARG00000044093	rpl13a	97	84.422	Danio_rerio
ENSETEG00000005732	-	100	94.581	ENSDNOG00000035370	-	100	94.581	Dasypus_novemcinctus
ENSETEG00000005732	-	100	94.089	ENSDORG00000012194	-	100	94.089	Dipodomys_ordii
ENSETEG00000005732	-	97	54.592	FBgn0037351	RpL13A	96	54.592	Drosophila_melanogaster
ENSETEG00000005732	-	98	66.332	ENSEBUG00000007234	rpl13a	100	65.426	Eptatretus_burgeri
ENSETEG00000005732	-	98	94.444	ENSEASG00005005007	RPL13A	86	94.444	Equus_asinus_asinus
ENSETEG00000005732	-	100	94.089	ENSECAG00000015832	RPL13A	100	94.089	Equus_caballus
ENSETEG00000005732	-	74	92.715	ENSEEUG00000014071	RPL13A	74	92.715	Erinaceus_europaeus
ENSETEG00000005732	-	98	83.920	ENSELUG00000014577	rpl13a	97	83.920	Esox_lucius
ENSETEG00000005732	-	100	94.581	ENSFCAG00000009194	RPL13A	100	94.581	Felis_catus
ENSETEG00000005732	-	100	94.089	ENSFDAG00000016892	RPL13A	100	94.089	Fukomys_damarensis
ENSETEG00000005732	-	98	84.925	ENSFHEG00000004165	rpl13a	97	84.925	Fundulus_heteroclitus
ENSETEG00000005732	-	98	82.915	ENSGMOG00000010791	rpl13a	93	82.915	Gadus_morhua
ENSETEG00000005732	-	98	84.422	ENSGAFG00000007578	rpl13a	97	84.422	Gambusia_affinis
ENSETEG00000005732	-	98	84.422	ENSGACG00000009310	rpl13a	99	83.920	Gasterosteus_aculeatus
ENSETEG00000005732	-	94	80.208	ENSGAGG00000002437	RPL13A	74	80.208	Gopherus_agassizii
ENSETEG00000005732	-	100	94.581	ENSGGOG00000002932	RPL13A	100	94.581	Gorilla_gorilla
ENSETEG00000005732	-	67	89.706	ENSGGOG00000039904	-	72	89.706	Gorilla_gorilla
ENSETEG00000005732	-	100	92.118	ENSGGOG00000023324	-	100	92.118	Gorilla_gorilla
ENSETEG00000005732	-	68	90.647	ENSGGOG00000039250	-	84	90.647	Gorilla_gorilla
ENSETEG00000005732	-	98	84.925	ENSHBUG00000023575	rpl13a	97	84.925	Haplochromis_burtoni
ENSETEG00000005732	-	100	94.089	ENSHGLG00000010070	RPL13A	100	94.472	Heterocephalus_glaber_female
ENSETEG00000005732	-	100	94.089	ENSHGLG00100010359	RPL13A	100	94.089	Heterocephalus_glaber_male
ENSETEG00000005732	-	98	84.343	ENSHCOG00000007902	rpl13a	99	84.343	Hippocampus_comes
ENSETEG00000005732	-	98	83.417	ENSIPUG00000014098	rpl13a	94	83.417	Ictalurus_punctatus
ENSETEG00000005732	-	100	92.157	ENSSTOG00000021558	RPL13A	99	93.939	Ictidomys_tridecemlineatus
ENSETEG00000005732	-	100	94.581	ENSJJAG00000000343	-	100	94.581	Jaculus_jaculus
ENSETEG00000005732	-	98	83.417	ENSKMAG00000009039	rpl13a	97	83.417	Kryptolebias_marmoratus
ENSETEG00000005732	-	98	84.925	ENSLBEG00000005434	rpl13a	97	84.925	Labrus_bergylta
ENSETEG00000005732	-	97	80.905	ENSLACG00000019092	RPL13A	95	81.313	Latimeria_chalumnae
ENSETEG00000005732	-	98	85.427	ENSLOCG00000004388	rpl13a	97	85.427	Lepisosteus_oculatus
ENSETEG00000005732	-	100	94.089	ENSLAFG00000016213	RPL13A	100	94.089	Loxodonta_africana
ENSETEG00000005732	-	100	84.236	ENSMFAG00000003121	-	100	84.729	Macaca_fascicularis
ENSETEG00000005732	-	100	94.089	ENSMFAG00000001097	-	100	94.089	Macaca_fascicularis
ENSETEG00000005732	-	100	86.538	ENSMFAG00000002765	-	100	86.538	Macaca_fascicularis
ENSETEG00000005732	-	100	87.019	ENSMMUG00000009018	-	100	87.019	Macaca_mulatta
ENSETEG00000005732	-	100	94.089	ENSMMUG00000007551	RPL13A	100	94.089	Macaca_mulatta
ENSETEG00000005732	-	100	94.089	ENSMNEG00000034977	RPL13A	100	94.089	Macaca_nemestrina
ENSETEG00000005732	-	58	94.068	ENSMLEG00000038132	-	100	94.068	Mandrillus_leucophaeus
ENSETEG00000005732	-	98	84.925	ENSMAMG00000007425	rpl13a	84	84.925	Mastacembelus_armatus
ENSETEG00000005732	-	98	84.925	ENSMZEG00005004952	rpl13a	97	84.925	Maylandia_zebra
ENSETEG00000005732	-	100	80.296	ENSMAUG00000012558	-	100	80.296	Mesocricetus_auratus
ENSETEG00000005732	-	100	92.118	ENSMAUG00000016402	-	100	92.118	Mesocricetus_auratus
ENSETEG00000005732	-	100	87.192	ENSMAUG00000020351	-	100	87.192	Mesocricetus_auratus
ENSETEG00000005732	-	100	76.355	ENSMICG00000043457	-	100	76.355	Microcebus_murinus
ENSETEG00000005732	-	100	95.074	ENSMICG00000009318	-	100	95.074	Microcebus_murinus
ENSETEG00000005732	-	100	76.847	ENSMOCG00000016427	-	100	76.847	Microtus_ochrogaster
ENSETEG00000005732	-	100	92.118	ENSMOCG00000002774	-	100	92.118	Microtus_ochrogaster
ENSETEG00000005732	-	98	85.427	ENSMMOG00000010248	rpl13a	98	85.427	Mola_mola
ENSETEG00000005732	-	98	84.925	ENSMALG00000012804	rpl13a	97	84.925	Monopterus_albus
ENSETEG00000005732	-	100	91.133	MGP_CAROLIEiJ_G0029698	-	100	91.133	Mus_caroli
ENSETEG00000005732	-	100	91.626	ENSMUSG00000074129	Rpl13a	100	91.626	Mus_musculus
ENSETEG00000005732	-	100	91.133	MGP_PahariEiJ_G0012923	-	100	91.133	Mus_pahari
ENSETEG00000005732	-	100	91.626	MGP_SPRETEiJ_G0030796	Rpl13a	100	91.626	Mus_spretus
ENSETEG00000005732	-	100	88.670	MGP_SPRETEiJ_G0017763	-	100	88.670	Mus_spretus
ENSETEG00000005732	-	100	94.581	ENSMPUG00000002604	RPL13A	100	94.581	Mustela_putorius_furo
ENSETEG00000005732	-	98	95.477	ENSMLUG00000004945	RPL13A	98	95.477	Myotis_lucifugus
ENSETEG00000005732	-	100	93.596	ENSNGAG00000021984	-	100	93.596	Nannospalax_galili
ENSETEG00000005732	-	98	85.354	ENSNBRG00000024166	rpl13a	97	84.925	Neolamprologus_brichardi
ENSETEG00000005732	-	99	88.557	ENSNLEG00000012957	-	97	88.557	Nomascus_leucogenys
ENSETEG00000005732	-	97	91.878	ENSNLEG00000033213	-	98	91.878	Nomascus_leucogenys
ENSETEG00000005732	-	100	86.408	ENSNLEG00000005211	RPL13A	100	86.408	Nomascus_leucogenys
ENSETEG00000005732	-	85	80.347	ENSMEUG00000005199	RPL13A	100	80.347	Notamacropus_eugenii
ENSETEG00000005732	-	100	89.163	ENSOPRG00000001194	RPL13A	100	89.163	Ochotona_princeps
ENSETEG00000005732	-	100	93.103	ENSODEG00000019218	-	100	93.103	Octodon_degus
ENSETEG00000005732	-	100	94.089	ENSODEG00000016831	-	100	94.089	Octodon_degus
ENSETEG00000005732	-	100	92.611	ENSODEG00000020233	-	100	92.611	Octodon_degus
ENSETEG00000005732	-	100	93.596	ENSODEG00000020468	-	100	93.596	Octodon_degus
ENSETEG00000005732	-	98	84.422	ENSONIG00000003560	rpl13a	97	84.422	Oreochromis_niloticus
ENSETEG00000005732	-	100	89.655	ENSOANG00000000828	RPL13A	100	89.655	Ornithorhynchus_anatinus
ENSETEG00000005732	-	77	82.803	ENSOCUG00000026592	-	97	82.803	Oryctolagus_cuniculus
ENSETEG00000005732	-	98	83.920	ENSORLG00000014599	rpl13a	87	83.920	Oryzias_latipes
ENSETEG00000005732	-	98	83.920	ENSORLG00020013090	rpl13a	87	83.920	Oryzias_latipes_hni
ENSETEG00000005732	-	99	83.582	ENSORLG00015011634	rpl13a	98	83.582	Oryzias_latipes_hsok
ENSETEG00000005732	-	98	84.925	ENSOMEG00000010232	rpl13a	87	84.925	Oryzias_melastigma
ENSETEG00000005732	-	99	92.611	ENSOGAG00000030280	-	99	92.611	Otolemur_garnettii
ENSETEG00000005732	-	100	94.581	ENSOGAG00000004625	-	100	94.581	Otolemur_garnettii
ENSETEG00000005732	-	100	88.670	ENSOGAG00000026177	-	100	88.670	Otolemur_garnettii
ENSETEG00000005732	-	97	76.020	ENSOARG00000017798	-	96	76.020	Ovis_aries
ENSETEG00000005732	-	100	92.118	ENSOARG00000020159	-	100	92.118	Ovis_aries
ENSETEG00000005732	-	99	93.532	ENSOARG00000013169	-	99	93.532	Ovis_aries
ENSETEG00000005732	-	67	89.706	ENSPPAG00000006023	-	71	89.706	Pan_paniscus
ENSETEG00000005732	-	100	94.581	ENSPPAG00000037365	RPL13A	100	94.581	Pan_paniscus
ENSETEG00000005732	-	100	91.626	ENSPPAG00000023659	-	100	91.626	Pan_paniscus
ENSETEG00000005732	-	100	91.626	ENSPPAG00000019107	-	100	91.626	Pan_paniscus
ENSETEG00000005732	-	100	94.581	ENSPPRG00000011265	RPL13A	100	94.581	Panthera_pardus
ENSETEG00000005732	-	70	93.662	ENSPTRG00000042978	-	100	93.662	Pan_troglodytes
ENSETEG00000005732	-	100	91.133	ENSPTRG00000049426	-	100	91.133	Pan_troglodytes
ENSETEG00000005732	-	100	94.581	ENSPTRG00000011298	RPL13A	100	94.581	Pan_troglodytes
ENSETEG00000005732	-	67	94.815	ENSPTRG00000052284	-	92	94.815	Pan_troglodytes
ENSETEG00000005732	-	100	93.103	ENSPTRG00000050666	-	100	93.103	Pan_troglodytes
ENSETEG00000005732	-	100	94.089	ENSPANG00000012050	RPL13A	100	94.089	Papio_anubis
ENSETEG00000005732	-	98	83.920	ENSPKIG00000000720	rpl13a	97	83.920	Paramormyrops_kingsleyae
ENSETEG00000005732	-	98	86.935	ENSPSIG00000018234	RPL13A	100	86.935	Pelodiscus_sinensis
ENSETEG00000005732	-	98	84.925	ENSPMGG00000010552	rpl13a	96	84.925	Periophthalmus_magnuspinnatus
ENSETEG00000005732	-	100	92.611	ENSPEMG00000009221	-	100	92.611	Peromyscus_maniculatus_bairdii
ENSETEG00000005732	-	98	82.915	ENSPMAG00000006362	rpl13a	97	82.915	Petromyzon_marinus
ENSETEG00000005732	-	83	75.148	ENSPCIG00000014063	-	90	75.148	Phascolarctos_cinereus
ENSETEG00000005732	-	100	90.640	ENSPCIG00000007340	-	100	90.640	Phascolarctos_cinereus
ENSETEG00000005732	-	98	83.920	ENSPFOG00000013546	rpl13a	97	83.920	Poecilia_formosa
ENSETEG00000005732	-	98	83.920	ENSPLAG00000016066	rpl13a	97	83.920	Poecilia_latipinna
ENSETEG00000005732	-	98	83.920	ENSPMEG00000018384	rpl13a	97	83.920	Poecilia_mexicana
ENSETEG00000005732	-	98	83.920	ENSPREG00000020079	rpl13a	97	83.920	Poecilia_reticulata
ENSETEG00000005732	-	100	94.581	ENSPPYG00000006798	-	100	94.581	Pongo_abelii
ENSETEG00000005732	-	100	94.581	ENSPPYG00000010251	-	100	94.581	Pongo_abelii
ENSETEG00000005732	-	100	94.581	ENSPCAG00000004086	RPL13A	100	94.581	Procavia_capensis
ENSETEG00000005732	-	100	95.074	ENSPCOG00000023898	RPL13A	100	95.074	Propithecus_coquereli
ENSETEG00000005732	-	98	77.833	ENSPCOG00000016293	-	98	77.833	Propithecus_coquereli
ENSETEG00000005732	-	100	94.089	ENSPVAG00000010958	RPL13A	100	94.089	Pteropus_vampyrus
ENSETEG00000005732	-	98	84.925	ENSPNYG00000019779	rpl13a	97	84.925	Pundamilia_nyererei
ENSETEG00000005732	-	98	83.838	ENSPNAG00000026887	rpl13a	97	83.838	Pygocentrus_nattereri
ENSETEG00000005732	-	100	92.611	ENSRNOG00000062298	Rpl13a	100	92.611	Rattus_norvegicus
ENSETEG00000005732	-	100	93.103	ENSRBIG00000044737	RPL13A	99	93.103	Rhinopithecus_bieti
ENSETEG00000005732	-	67	88.235	ENSRROG00000028369	-	68	88.235	Rhinopithecus_roxellana
ENSETEG00000005732	-	100	94.089	ENSRROG00000042976	-	100	94.089	Rhinopithecus_roxellana
ENSETEG00000005732	-	100	88.670	ENSRROG00000036041	-	100	88.670	Rhinopithecus_roxellana
ENSETEG00000005732	-	84	59.064	YNL069C	RPL16B	86	59.064	Saccharomyces_cerevisiae
ENSETEG00000005732	-	84	60.819	YIL133C	RPL16A	86	60.819	Saccharomyces_cerevisiae
ENSETEG00000005732	-	100	76.364	ENSSBOG00000024635	-	100	76.364	Saimiri_boliviensis_boliviensis
ENSETEG00000005732	-	100	93.596	ENSSBOG00000028481	RPL13A	100	93.970	Saimiri_boliviensis_boliviensis
ENSETEG00000005732	-	98	81.910	ENSSFOG00015001972	rpl13a	97	81.910	Scleropages_formosus
ENSETEG00000005732	-	98	84.422	ENSSDUG00000017181	rpl13a	100	84.422	Seriola_dumerili
ENSETEG00000005732	-	81	59.146	ENSSLDG00000007920	-	85	58.929	Seriola_lalandi_dorsalis
ENSETEG00000005732	-	98	84.422	ENSSLDG00000002561	rpl13a	97	84.422	Seriola_lalandi_dorsalis
ENSETEG00000005732	-	53	69.725	ENSSPUG00000014548	RPL13A	86	69.725	Sphenodon_punctatus
ENSETEG00000005732	-	95	70.833	ENSSSCG00000031204	-	92	76.163	Sus_scrofa
ENSETEG00000005732	-	100	93.137	ENSSSCG00000034448	-	100	93.137	Sus_scrofa
ENSETEG00000005732	-	98	84.422	ENSTRUG00000023611	rpl13a	97	84.422	Takifugu_rubripes
ENSETEG00000005732	-	100	93.596	ENSTTRG00000005485	RPL13A	100	93.596	Tursiops_truncatus
ENSETEG00000005732	-	100	95.074	ENSUAMG00000008438	RPL13A	100	95.074	Ursus_americanus
ENSETEG00000005732	-	100	95.074	ENSUMAG00000002385	RPL13A	100	95.074	Ursus_maritimus
ENSETEG00000005732	-	98	91.919	ENSVPAG00000006564	RPL13A	100	91.919	Vicugna_pacos
ENSETEG00000005732	-	100	95.074	ENSVVUG00000022597	RPL13A	100	95.074	Vulpes_vulpes
ENSETEG00000005732	-	98	87.940	ENSXETG00000014144	rpl13a	86	87.940	Xenopus_tropicalis
ENSETEG00000005732	-	98	84.925	ENSXCOG00000004826	rpl13a	97	84.925	Xiphophorus_couchianus
ENSETEG00000005732	-	98	84.925	ENSXMAG00000027604	rpl13a	97	84.925	Xiphophorus_maculatus