EuRBPDB

Description

RBDs

Transcripts

Gene Model

PPI

Paralogs

Orthologs

Description

Ensembl ID: ENSGACG00000020055 (Gene tree)
Gene ID: -
Gene Symbol: snrpf
Alias: snrpfl
Full Name: small nuclear ribonucleoprotein polypeptide F
Gene Type: protein_coding
Species: Gasterosteus_aculeatus
Status: putative
Strand: Minus strand
Length: 1033 bases
Position: chrgroupIV:32244029-32245061
Accession: ZDB-GENE-040930-9
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
ENSGACP00000026497	LSM	PF01423.22	1.5e-20	1	1

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
ENSGACT00000026548	-	258	-	ENSGACP00000026497	85 (aa)	-	G3Q9D2

Gene Model

Click here to download ENSGACG00000020055's gene model file

Protein-Protein Interaction (PPI)

Clik here to download ENSGACG00000020055's network

Paralogs

Ensembl ID	Gene Symbol	Coverage	Identiy	Paralog	Gene Symbol	Coverage	Identiy
ENSGACG00000020055	snrpf	78	50.000	ENSGACG00000017223	lsm6	83	50.000

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
ENSGACG00000020055	snrpf	100	96.471	ENSG00000139343	SNRPF	99	96.471	Homo_sapiens
ENSGACG00000020055	snrpf	100	98.824	ENSAPOG00000003377	snrpf	66	98.824	Acanthochromis_polyacanthus
ENSGACG00000020055	snrpf	100	96.471	ENSAMEG00000002166	-	100	96.471	Ailuropoda_melanoleuca
ENSGACG00000020055	snrpf	88	82.667	ENSAMEG00000018718	-	86	83.784	Ailuropoda_melanoleuca
ENSGACG00000020055	snrpf	99	98.810	ENSACIG00000013216	snrpf	99	98.810	Amphilophus_citrinellus
ENSGACG00000020055	snrpf	99	98.810	ENSAOCG00000015108	snrpf	99	98.810	Amphiprion_ocellaris
ENSGACG00000020055	snrpf	99	98.810	ENSAPEG00000022022	snrpf	99	98.810	Amphiprion_percula
ENSGACG00000020055	snrpf	99	98.810	ENSATEG00000022625	snrpf	90	98.810	Anabas_testudineus
ENSGACG00000020055	snrpf	87	94.595	ENSACAG00000014073	SNRPF	85	94.595	Anolis_carolinensis
ENSGACG00000020055	snrpf	84	90.141	ENSANAG00000020297	-	65	90.141	Aotus_nancymaae
ENSGACG00000020055	snrpf	100	96.471	ENSANAG00000032845	SNRPF	99	96.471	Aotus_nancymaae
ENSGACG00000020055	snrpf	87	66.216	ENSANAG00000025860	-	83	66.216	Aotus_nancymaae
ENSGACG00000020055	snrpf	99	98.810	ENSACLG00000016066	snrpf	99	98.810	Astatotilapia_calliptera
ENSGACG00000020055	snrpf	100	98.824	ENSAMXG00000007421	snrpf	99	98.824	Astyanax_mexicanus
ENSGACG00000020055	snrpf	91	90.909	ENSBTAG00000050012	-	91	90.541	Bos_taurus
ENSGACG00000020055	snrpf	100	96.471	ENSBTAG00000016271	SNRPF	99	96.471	Bos_taurus
ENSGACG00000020055	snrpf	91	70.130	WBGene00004918	snr-5	85	72.222	Caenorhabditis_elegans
ENSGACG00000020055	snrpf	100	96.471	ENSCJAG00000002705	-	99	96.471	Callithrix_jacchus
ENSGACG00000020055	snrpf	84	90.141	ENSCAFG00000031151	-	85	90.411	Canis_familiaris
ENSGACG00000020055	snrpf	100	96.471	ENSCAFG00000006395	-	77	96.471	Canis_familiaris
ENSGACG00000020055	snrpf	84	90.141	ENSCAFG00020010823	-	85	90.411	Canis_lupus_dingo
ENSGACG00000020055	snrpf	92	84.615	ENSCAFG00020013161	-	74	92.857	Canis_lupus_dingo
ENSGACG00000020055	snrpf	95	90.123	ENSTSYG00000031636	-	61	91.781	Carlito_syrichta
ENSGACG00000020055	snrpf	87	82.432	ENSTSYG00000030635	-	86	82.432	Carlito_syrichta
ENSGACG00000020055	snrpf	87	90.667	ENSTSYG00000036502	-	86	90.667	Carlito_syrichta
ENSGACG00000020055	snrpf	100	96.471	ENSCAPG00000007242	SNRPF	100	96.471	Cavia_aperea
ENSGACG00000020055	snrpf	100	96.471	ENSCPOG00000013898	SNRPF	99	96.471	Cavia_porcellus
ENSGACG00000020055	snrpf	100	95.294	ENSCCAG00000017356	-	86	95.946	Cebus_capucinus
ENSGACG00000020055	snrpf	99	86.905	ENSCCAG00000033627	-	57	87.671	Cebus_capucinus
ENSGACG00000020055	snrpf	100	96.471	ENSCCAG00000037378	-	99	96.471	Cebus_capucinus
ENSGACG00000020055	snrpf	100	96.471	ENSCLAG00000007200	SNRPF	99	96.471	Chinchilla_lanigera
ENSGACG00000020055	snrpf	100	96.471	ENSCSAG00000003602	-	69	96.471	Chlorocebus_sabaeus
ENSGACG00000020055	snrpf	74	95.238	ENSCHOG00000009392	SNRPF	98	95.238	Choloepus_hoffmanni
ENSGACG00000020055	snrpf	100	84.706	ENSCPBG00000001985	-	88	86.486	Chrysemys_picta_bellii
ENSGACG00000020055	snrpf	87	87.838	ENSCPBG00000006227	-	86	87.838	Chrysemys_picta_bellii
ENSGACG00000020055	snrpf	100	87.059	ENSCPBG00000026463	-	86	87.838	Chrysemys_picta_bellii
ENSGACG00000020055	snrpf	100	90.588	ENSCPBG00000024287	-	60	91.892	Chrysemys_picta_bellii
ENSGACG00000020055	snrpf	100	85.882	ENSCSAVG00000007498	-	84	87.838	Ciona_savignyi
ENSGACG00000020055	snrpf	100	96.471	ENSCGRG00001017100	-	99	96.471	Cricetulus_griseus_chok1gshd
ENSGACG00000020055	snrpf	86	89.041	ENSCGRG00001008965	-	57	89.041	Cricetulus_griseus_chok1gshd
ENSGACG00000020055	snrpf	92	89.744	ENSCGRG00000018878	-	91	89.041	Cricetulus_griseus_crigri
ENSGACG00000020055	snrpf	100	96.471	ENSCGRG00000000336	-	99	96.471	Cricetulus_griseus_crigri
ENSGACG00000020055	snrpf	100	97.647	ENSCSEG00000004748	snrpf	99	97.647	Cynoglossus_semilaevis
ENSGACG00000020055	snrpf	100	98.824	ENSCVAG00000000397	snrpf	67	98.824	Cyprinodon_variegatus
ENSGACG00000020055	snrpf	100	96.552	ENSDARG00000105037	snrpf	99	96.552	Danio_rerio
ENSGACG00000020055	snrpf	100	96.471	ENSDNOG00000043775	-	54	96.471	Dasypus_novemcinctus
ENSGACG00000020055	snrpf	87	94.595	ENSDNOG00000048989	-	66	94.595	Dasypus_novemcinctus
ENSGACG00000020055	snrpf	87	79.730	ENSDNOG00000049373	-	64	79.730	Dasypus_novemcinctus
ENSGACG00000020055	snrpf	84	87.324	ENSDORG00000025351	-	99	87.324	Dipodomys_ordii
ENSGACG00000020055	snrpf	69	93.220	ENSDORG00000029740	-	72	93.220	Dipodomys_ordii
ENSGACG00000020055	snrpf	99	76.190	FBgn0000426	SmF	83	79.452	Drosophila_melanogaster
ENSGACG00000020055	snrpf	100	96.471	ENSETEG00000001558	-	99	96.471	Echinops_telfairi
ENSGACG00000020055	snrpf	96	74.390	ENSEBUG00000001793	-	72	85.075	Eptatretus_burgeri
ENSGACG00000020055	snrpf	100	96.471	ENSECAG00000019817	SNRPF	69	96.471	Equus_caballus
ENSGACG00000020055	snrpf	100	96.471	ENSEEUG00000013509	SNRPF	100	96.471	Erinaceus_europaeus
ENSGACG00000020055	snrpf	100	98.824	ENSELUG00000003121	RUXF	99	98.824	Esox_lucius
ENSGACG00000020055	snrpf	100	95.294	ENSELUG00000007737	RUXF	85	97.297	Esox_lucius
ENSGACG00000020055	snrpf	100	96.471	ENSFCAG00000029966	-	99	96.471	Felis_catus
ENSGACG00000020055	snrpf	98	75.904	ENSFCAG00000031148	-	71	74.026	Felis_catus
ENSGACG00000020055	snrpf	87	72.973	ENSFCAG00000021942	-	85	72.973	Felis_catus
ENSGACG00000020055	snrpf	100	96.471	ENSFALG00000010799	SNRPF	99	96.471	Ficedula_albicollis
ENSGACG00000020055	snrpf	100	96.471	ENSFDAG00000002831	-	99	96.471	Fukomys_damarensis
ENSGACG00000020055	snrpf	100	98.824	ENSFHEG00000003240	snrpf	75	98.824	Fundulus_heteroclitus
ENSGACG00000020055	snrpf	100	98.824	ENSGMOG00000007741	snrpf	100	98.824	Gadus_morhua
ENSGACG00000020055	snrpf	100	96.471	ENSGALG00000011409	SNRPF	99	96.471	Gallus_gallus
ENSGACG00000020055	snrpf	99	98.810	ENSGAFG00000016187	snrpf	97	98.810	Gambusia_affinis
ENSGACG00000020055	snrpf	87	89.189	ENSGAGG00000012945	-	68	90.541	Gopherus_agassizii
ENSGACG00000020055	snrpf	93	88.608	ENSGAGG00000022991	-	80	91.176	Gopherus_agassizii
ENSGACG00000020055	snrpf	71	86.667	ENSGAGG00000011163	-	86	89.796	Gopherus_agassizii
ENSGACG00000020055	snrpf	80	82.353	ENSGAGG00000014132	-	82	82.353	Gopherus_agassizii
ENSGACG00000020055	snrpf	74	92.063	ENSGAGG00000020974	-	53	92.063	Gopherus_agassizii
ENSGACG00000020055	snrpf	100	96.471	ENSGGOG00000001127	-	99	96.471	Gorilla_gorilla
ENSGACG00000020055	snrpf	92	73.077	ENSGGOG00000039564	-	89	71.622	Gorilla_gorilla
ENSGACG00000020055	snrpf	98	77.108	ENSGGOG00000039936	-	82	74.324	Gorilla_gorilla
ENSGACG00000020055	snrpf	87	94.595	ENSHGLG00000021172	-	86	94.595	Heterocephalus_glaber_female
ENSGACG00000020055	snrpf	100	96.471	ENSHGLG00000009108	SNRPF	99	96.471	Heterocephalus_glaber_female
ENSGACG00000020055	snrpf	87	94.595	ENSHGLG00100019596	-	86	94.595	Heterocephalus_glaber_male
ENSGACG00000020055	snrpf	100	96.471	ENSHGLG00100008103	-	100	96.471	Heterocephalus_glaber_male
ENSGACG00000020055	snrpf	74	87.143	ENSHCOG00000018663	snrpf	55	87.143	Hippocampus_comes
ENSGACG00000020055	snrpf	100	98.824	ENSIPUG00000018994	snrpf	99	98.824	Ictalurus_punctatus
ENSGACG00000020055	snrpf	100	96.471	ENSSTOG00000010434	-	99	96.471	Ictidomys_tridecemlineatus
ENSGACG00000020055	snrpf	100	81.176	ENSJJAG00000010502	-	88	82.432	Jaculus_jaculus
ENSGACG00000020055	snrpf	100	77.647	ENSJJAG00000004558	-	88	78.378	Jaculus_jaculus
ENSGACG00000020055	snrpf	79	89.552	ENSJJAG00000004828	-	86	91.071	Jaculus_jaculus
ENSGACG00000020055	snrpf	100	96.471	ENSJJAG00000011272	-	99	96.471	Jaculus_jaculus
ENSGACG00000020055	snrpf	99	98.810	ENSKMAG00000013566	snrpf	98	98.810	Kryptolebias_marmoratus
ENSGACG00000020055	snrpf	100	97.647	ENSLBEG00000008800	snrpf	99	97.647	Labrus_bergylta
ENSGACG00000020055	snrpf	100	96.471	ENSLACG00000002456	-	73	96.471	Latimeria_chalumnae
ENSGACG00000020055	snrpf	100	98.824	ENSLOCG00000015744	snrpf	99	98.824	Lepisosteus_oculatus
ENSGACG00000020055	snrpf	100	96.471	ENSLAFG00000017426	SNRPF	99	96.471	Loxodonta_africana
ENSGACG00000020055	snrpf	99	97.619	ENSMAMG00000016593	snrpf	99	97.619	Mastacembelus_armatus
ENSGACG00000020055	snrpf	99	98.810	ENSMZEG00005014701	snrpf	99	98.810	Maylandia_zebra
ENSGACG00000020055	snrpf	99	96.429	ENSMGAG00000011453	-	76	96.429	Meleagris_gallopavo
ENSGACG00000020055	snrpf	74	95.238	ENSMAUG00000016909	-	85	95.238	Mesocricetus_auratus
ENSGACG00000020055	snrpf	74	95.238	ENSMAUG00000019737	-	85	95.238	Mesocricetus_auratus
ENSGACG00000020055	snrpf	72	88.525	ENSMAUG00000010289	-	66	92.000	Mesocricetus_auratus
ENSGACG00000020055	snrpf	87	91.892	ENSMAUG00000008799	-	86	91.892	Mesocricetus_auratus
ENSGACG00000020055	snrpf	87	89.189	ENSMICG00000030494	-	59	89.189	Microcebus_murinus
ENSGACG00000020055	snrpf	100	96.471	ENSMICG00000012153	-	99	96.471	Microcebus_murinus
ENSGACG00000020055	snrpf	80	79.412	ENSMOCG00000020220	-	87	79.412	Microtus_ochrogaster
ENSGACG00000020055	snrpf	87	82.432	ENSMOCG00000022166	-	81	82.432	Microtus_ochrogaster
ENSGACG00000020055	snrpf	100	96.471	ENSMOCG00000020437	Snrpf	99	96.471	Microtus_ochrogaster
ENSGACG00000020055	snrpf	100	97.647	ENSMMOG00000007127	snrpf	100	97.647	Mola_mola
ENSGACG00000020055	snrpf	100	96.471	ENSMODG00000008091	SNRPF	75	96.471	Monodelphis_domestica
ENSGACG00000020055	snrpf	99	98.810	ENSMALG00000012328	snrpf	99	98.810	Monopterus_albus
ENSGACG00000020055	snrpf	100	96.471	MGP_CAROLIEiJ_G0015735	-	99	96.471	Mus_caroli
ENSGACG00000020055	snrpf	100	96.471	ENSMUSG00000020018	Snrpf	99	96.471	Mus_musculus
ENSGACG00000020055	snrpf	100	96.471	MGP_PahariEiJ_G0031350	Snrpf	99	96.471	Mus_pahari
ENSGACG00000020055	snrpf	100	96.471	MGP_SPRETEiJ_G0016553	Snrpf	99	96.471	Mus_spretus
ENSGACG00000020055	snrpf	100	96.471	ENSMPUG00000007176	-	76	96.471	Mustela_putorius_furo
ENSGACG00000020055	snrpf	100	96.471	ENSMLUG00000027932	-	80	96.471	Myotis_lucifugus
ENSGACG00000020055	snrpf	100	96.471	ENSMLUG00000029201	-	80	96.471	Myotis_lucifugus
ENSGACG00000020055	snrpf	100	96.471	ENSMLUG00000026149	-	80	96.471	Myotis_lucifugus
ENSGACG00000020055	snrpf	100	96.471	ENSMLUG00000013577	-	100	96.471	Myotis_lucifugus
ENSGACG00000020055	snrpf	100	77.647	ENSMLUG00000026269	-	70	75.676	Myotis_lucifugus
ENSGACG00000020055	snrpf	74	88.889	ENSNGAG00000005851	-	71	88.333	Nannospalax_galili
ENSGACG00000020055	snrpf	100	96.471	ENSNGAG00000024292	-	99	96.471	Nannospalax_galili
ENSGACG00000020055	snrpf	100	98.824	ENSNBRG00000011682	snrpf	87	98.824	Neolamprologus_brichardi
ENSGACG00000020055	snrpf	87	70.270	ENSMEUG00000000671	SNRPF	87	70.270	Notamacropus_eugenii
ENSGACG00000020055	snrpf	100	96.471	ENSODEG00000010328	-	99	96.471	Octodon_degus
ENSGACG00000020055	snrpf	100	96.471	ENSOCUG00000010925	SNRPF	75	96.471	Oryctolagus_cuniculus
ENSGACG00000020055	snrpf	100	98.824	ENSORLG00020008216	snrpf	100	98.824	Oryzias_latipes_hni
ENSGACG00000020055	snrpf	100	98.824	ENSORLG00015007418	snrpf	100	98.824	Oryzias_latipes_hsok
ENSGACG00000020055	snrpf	99	100.000	ENSOMEG00000003018	snrpf	99	100.000	Oryzias_melastigma
ENSGACG00000020055	snrpf	87	94.595	ENSOGAG00000030323	-	86	94.595	Otolemur_garnettii
ENSGACG00000020055	snrpf	100	96.471	ENSOARG00000011853	-	74	96.471	Ovis_aries
ENSGACG00000020055	snrpf	98	77.108	ENSPPAG00000037544	-	82	74.324	Pan_paniscus
ENSGACG00000020055	snrpf	92	73.077	ENSPPAG00000037479	-	89	71.622	Pan_paniscus
ENSGACG00000020055	snrpf	100	96.471	ENSPPRG00000005875	-	99	96.471	Panthera_pardus
ENSGACG00000020055	snrpf	98	74.699	ENSPPRG00000007449	-	76	74.324	Panthera_pardus
ENSGACG00000020055	snrpf	100	96.471	ENSPTIG00000007520	SNRPF	99	96.471	Panthera_tigris_altaica
ENSGACG00000020055	snrpf	98	77.108	ENSPTIG00000005117	-	88	74.324	Panthera_tigris_altaica
ENSGACG00000020055	snrpf	92	73.077	ENSPTRG00000047717	-	89	71.622	Pan_troglodytes
ENSGACG00000020055	snrpf	98	77.108	ENSPTRG00000043814	-	82	74.324	Pan_troglodytes
ENSGACG00000020055	snrpf	100	98.824	ENSPKIG00000014784	snrpf	93	98.824	Paramormyrops_kingsleyae
ENSGACG00000020055	snrpf	100	96.471	ENSPSIG00000016462	SNRPF	99	96.471	Pelodiscus_sinensis
ENSGACG00000020055	snrpf	100	97.647	ENSPMGG00000023548	snrpf	100	97.647	Periophthalmus_magnuspinnatus
ENSGACG00000020055	snrpf	100	96.471	ENSPEMG00000023326	Snrpf	99	96.471	Peromyscus_maniculatus_bairdii
ENSGACG00000020055	snrpf	99	98.810	ENSPLAG00000023932	snrpf	99	98.810	Poecilia_latipinna
ENSGACG00000020055	snrpf	99	98.810	ENSPMEG00000003377	snrpf	99	98.810	Poecilia_mexicana
ENSGACG00000020055	snrpf	99	98.810	ENSPREG00000022635	snrpf	97	98.810	Poecilia_reticulata
ENSGACG00000020055	snrpf	100	96.471	ENSPPYG00000004847	-	99	96.471	Pongo_abelii
ENSGACG00000020055	snrpf	100	96.471	ENSPCOG00000023833	-	99	96.471	Propithecus_coquereli
ENSGACG00000020055	snrpf	100	96.471	ENSPVAG00000009036	SNRPF	99	96.471	Pteropus_vampyrus
ENSGACG00000020055	snrpf	99	98.810	ENSPNYG00000021712	snrpf	99	98.810	Pundamilia_nyererei
ENSGACG00000020055	snrpf	99	100.000	ENSPNAG00000019117	snrpf	99	100.000	Pygocentrus_nattereri
ENSGACG00000020055	snrpf	100	96.471	ENSRNOG00000005556	Snrpf	99	96.471	Rattus_norvegicus
ENSGACG00000020055	snrpf	87	93.243	ENSRBIG00000030414	-	86	93.243	Rhinopithecus_bieti
ENSGACG00000020055	snrpf	87	93.243	ENSRROG00000021975	-	86	93.243	Rhinopithecus_roxellana
ENSGACG00000020055	snrpf	82	59.155	YPR182W	SMX3	83	59.155	Saccharomyces_cerevisiae
ENSGACG00000020055	snrpf	100	96.471	ENSSBOG00000036179	-	99	96.471	Saimiri_boliviensis_boliviensis
ENSGACG00000020055	snrpf	87	81.081	ENSSBOG00000032297	-	85	81.081	Saimiri_boliviensis_boliviensis
ENSGACG00000020055	snrpf	87	95.946	ENSSBOG00000012868	-	86	95.946	Saimiri_boliviensis_boliviensis
ENSGACG00000020055	snrpf	91	76.250	ENSSBOG00000027638	-	90	76.250	Saimiri_boliviensis_boliviensis
ENSGACG00000020055	snrpf	100	96.471	ENSSHAG00000012294	SNRPF	77	96.471	Sarcophilus_harrisii
ENSGACG00000020055	snrpf	75	98.438	ENSSFOG00015021523	snrpf	96	98.438	Scleropages_formosus
ENSGACG00000020055	snrpf	100	100.000	ENSSMAG00000005533	snrpf	98	100.000	Scophthalmus_maximus
ENSGACG00000020055	snrpf	75	98.438	ENSSDUG00000016283	snrpf	76	98.438	Seriola_dumerili
ENSGACG00000020055	snrpf	94	70.000	ENSSLDG00000003885	-	90	72.973	Seriola_lalandi_dorsalis
ENSGACG00000020055	snrpf	99	98.810	ENSSLDG00000005119	snrpf	98	98.810	Seriola_lalandi_dorsalis
ENSGACG00000020055	snrpf	99	96.429	ENSSPUG00000014590	-	99	96.429	Sphenodon_punctatus
ENSGACG00000020055	snrpf	100	98.824	ENSSPAG00000003283	snrpf	97	98.824	Stegastes_partitus
ENSGACG00000020055	snrpf	100	96.471	ENSSSCG00000000895	SNRPF	99	96.471	Sus_scrofa
ENSGACG00000020055	snrpf	81	97.101	ENSTRUG00000014956	snrpf	100	97.101	Takifugu_rubripes
ENSGACG00000020055	snrpf	100	97.647	ENSTNIG00000012672	snrpf	99	97.647	Tetraodon_nigroviridis
ENSGACG00000020055	snrpf	87	70.270	ENSTTRG00000006070	SNRPF	86	70.270	Tursiops_truncatus
ENSGACG00000020055	snrpf	100	83.529	ENSUAMG00000001101	-	87	85.135	Ursus_americanus
ENSGACG00000020055	snrpf	100	96.471	ENSUAMG00000008176	-	99	96.471	Ursus_americanus
ENSGACG00000020055	snrpf	87	85.135	ENSUMAG00000006177	-	87	85.135	Ursus_maritimus
ENSGACG00000020055	snrpf	100	96.471	ENSUMAG00000003528	-	99	96.471	Ursus_maritimus
ENSGACG00000020055	snrpf	100	96.471	ENSVPAG00000008712	-	99	96.471	Vicugna_pacos
ENSGACG00000020055	snrpf	91	88.312	ENSVVUG00000023821	-	95	87.838	Vulpes_vulpes
ENSGACG00000020055	snrpf	87	75.000	ENSXETG00000021630	snrpf	87	75.000	Xenopus_tropicalis
ENSGACG00000020055	snrpf	99	98.810	ENSXCOG00000013860	snrpf	97	98.810	Xiphophorus_couchianus
ENSGACG00000020055	snrpf	99	98.810	ENSXMAG00000015448	snrpf	99	98.810	Xiphophorus_maculatus