EuRBPDB

Description

RBDs

Transcripts

Gene Model

PPI

Orthologs

Description

Ensembl ID: ENSGALG00000009677 (Gene tree)
Gene ID: 395268
Gene Symbol: SSB
Alias: UBP
Full Name: Gallus gallus Sjogren syndrome antigen B (SSB)%2C mRNA.
Gene Type: protein_coding
Species: Gallus_gallus
Status: putative
Strand: Minus strand
Length: 8765 bases
Position: chr7:18575885-18584649
Accession: NM_001193669
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
ENSGALP00000015735	RRM_1	PF00076.22	6.9e-14	1	1
ENSGALP00000062135	RRM_1	PF00076.22	6.9e-14	1	1
ENSGALP00000067291	RRM_1	PF00076.22	6.9e-14	1	1
ENSGALP00000070268	RRM_1	PF00076.22	6.9e-14	1	1
ENSGALP00000015735	RRM_3	PF08777.11	1.6e-27	1	1
ENSGALP00000062135	RRM_3	PF08777.11	1.6e-27	1	1
ENSGALP00000067291	RRM_3	PF08777.11	1.6e-27	1	1
ENSGALP00000070268	RRM_3	PF08777.11	1.6e-27	1	1
ENSGALP00000015735	La	PF05383.17	2.9e-21	1	1
ENSGALP00000062135	La	PF05383.17	2.9e-21	1	1
ENSGALP00000067291	La	PF05383.17	2.9e-21	1	1
ENSGALP00000070268	La	PF05383.17	2.9e-21	1	1

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
ENSGALT00000093425	-	2442	-	ENSGALP00000070268	404 (aa)	-	UPI0000ECB733
ENSGALT00000100882	-	1972	-	ENSGALP00000067291	404 (aa)	-	UPI0000ECB733
ENSGALT00000084239	-	1648	XM_015289615	ENSGALP00000062135	404 (aa)	XP_015145101	Q8QHI5
ENSGALT00000015751	-	1948	-	ENSGALP00000015735	404 (aa)	-	F6R1X6

Gene Model

Click here to download ENSGALG00000009677's gene model file

Protein-Protein Interaction (PPI)

Clik here to download ENSGALG00000009677's network

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
ENSGALG00000009677	SSB	100	74.265	ENSG00000138385	SSB	100	79.167	Homo_sapiens
ENSGALG00000009677	SSB	93	57.031	ENSAPOG00000003448	ssb	98	58.333	Acanthochromis_polyacanthus
ENSGALG00000009677	SSB	100	71.631	ENSAMEG00000010507	SSB	100	71.631	Ailuropoda_melanoleuca
ENSGALG00000009677	SSB	91	57.838	ENSACIG00000001738	ssb	97	59.140	Amphilophus_citrinellus
ENSGALG00000009677	SSB	93	57.031	ENSAOCG00000020398	ssb	98	58.333	Amphiprion_ocellaris
ENSGALG00000009677	SSB	93	57.292	ENSAPEG00000020395	ssb	98	58.594	Amphiprion_percula
ENSGALG00000009677	SSB	99	57.353	ENSATEG00000016884	ssb	99	58.578	Anabas_testudineus
ENSGALG00000009677	SSB	100	89.176	ENSAPLG00000006109	SSB	100	89.882	Anas_platyrhynchos
ENSGALG00000009677	SSB	99	70.370	ENSACAG00000017008	SSB	99	70.617	Anolis_carolinensis
ENSGALG00000009677	SSB	100	67.822	ENSANAG00000027195	-	100	68.889	Aotus_nancymaae
ENSGALG00000009677	SSB	99	54.229	ENSACLG00000010798	ssb	99	54.839	Astatotilapia_calliptera
ENSGALG00000009677	SSB	97	55.473	ENSAMXG00000012606	ssb	99	56.522	Astyanax_mexicanus
ENSGALG00000009677	SSB	100	75.245	ENSBTAG00000008716	SSB	100	75.245	Bos_taurus
ENSGALG00000009677	SSB	74	36.184	WBGene00016653	C44E4.4	92	34.771	Caenorhabditis_elegans
ENSGALG00000009677	SSB	100	74.510	ENSCJAG00000045073	SSB	100	74.510	Callithrix_jacchus
ENSGALG00000009677	SSB	100	75.061	ENSCAFG00000012418	SSB	100	75.061	Canis_familiaris
ENSGALG00000009677	SSB	100	75.061	ENSCAFG00020002900	SSB	100	75.061	Canis_lupus_dingo
ENSGALG00000009677	SSB	100	75.245	ENSCHIG00000022624	-	100	75.245	Capra_hircus
ENSGALG00000009677	SSB	100	71.324	ENSCHIG00000022196	-	100	71.324	Capra_hircus
ENSGALG00000009677	SSB	100	74.265	ENSTSYG00000000097	SSB	100	74.265	Carlito_syrichta
ENSGALG00000009677	SSB	100	76.528	ENSCPOG00000003338	SSB	100	76.528	Cavia_porcellus
ENSGALG00000009677	SSB	100	75.000	ENSCCAG00000035147	SSB	100	75.000	Cebus_capucinus
ENSGALG00000009677	SSB	100	54.902	ENSCATG00000042909	-	100	54.167	Cercocebus_atys
ENSGALG00000009677	SSB	78	67.674	ENSCATG00000036701	SSB	93	67.674	Cercocebus_atys
ENSGALG00000009677	SSB	100	75.735	ENSCLAG00000015617	-	100	75.735	Chinchilla_lanigera
ENSGALG00000009677	SSB	100	74.755	ENSCSAG00000015200	SSB	89	74.755	Chlorocebus_sabaeus
ENSGALG00000009677	SSB	98	68.750	ENSCSAG00000018390	-	100	68.750	Chlorocebus_sabaeus
ENSGALG00000009677	SSB	100	74.510	ENSCHOG00000007763	SSB	100	74.510	Choloepus_hoffmanni
ENSGALG00000009677	SSB	100	84.841	ENSCPBG00000006519	SSB	100	85.819	Chrysemys_picta_bellii
ENSGALG00000009677	SSB	99	65.594	ENSCANG00000033240	-	99	65.594	Colobus_angolensis_palliatus
ENSGALG00000009677	SSB	96	59.439	ENSCANG00000028838	-	93	59.439	Colobus_angolensis_palliatus
ENSGALG00000009677	SSB	100	74.755	ENSCANG00000027695	SSB	100	74.755	Colobus_angolensis_palliatus
ENSGALG00000009677	SSB	99	64.792	ENSCGRG00001024831	Ssb	99	66.748	Cricetulus_griseus_chok1gshd
ENSGALG00000009677	SSB	99	64.792	ENSCGRG00000005387	Ssb	99	66.748	Cricetulus_griseus_crigri
ENSGALG00000009677	SSB	97	55.138	ENSCSEG00000011599	ssb	99	56.359	Cynoglossus_semilaevis
ENSGALG00000009677	SSB	99	55.854	ENSCVAG00000021558	ssb	99	57.073	Cyprinodon_variegatus
ENSGALG00000009677	SSB	98	58.416	ENSDARG00000029252	ssb	99	59.012	Danio_rerio
ENSGALG00000009677	SSB	100	75.000	ENSDNOG00000013450	SSB	100	75.000	Dasypus_novemcinctus
ENSGALG00000009677	SSB	100	66.667	ENSDORG00000005543	Ssb	100	68.599	Dipodomys_ordii
ENSGALG00000009677	SSB	82	34.648	FBgn0011638	La	87	34.626	Drosophila_melanogaster
ENSGALG00000009677	SSB	50	76.617	ENSETEG00000004614	-	52	76.617	Echinops_telfairi
ENSGALG00000009677	SSB	91	47.745	ENSEBUG00000001237	ssb	94	47.090	Eptatretus_burgeri
ENSGALG00000009677	SSB	100	75.490	ENSEASG00005022835	SSB	100	75.490	Equus_asinus_asinus
ENSGALG00000009677	SSB	100	75.490	ENSECAG00000018495	SSB	100	75.490	Equus_caballus
ENSGALG00000009677	SSB	76	62.418	ENSEEUG00000008077	SSB	78	62.418	Erinaceus_europaeus
ENSGALG00000009677	SSB	99	61.916	ENSELUG00000020929	ssb	99	62.899	Esox_lucius
ENSGALG00000009677	SSB	100	74.328	ENSFCAG00000011601	SSB	100	74.328	Felis_catus
ENSGALG00000009677	SSB	100	93.350	ENSFALG00000000930	SSB	100	94.089	Ficedula_albicollis
ENSGALG00000009677	SSB	100	75.490	ENSFDAG00000000598	SSB	100	75.490	Fukomys_damarensis
ENSGALG00000009677	SSB	62	58.366	ENSFHEG00000003268	ssb	93	55.482	Fundulus_heteroclitus
ENSGALG00000009677	SSB	97	54.126	ENSGMOG00000002664	ssb	99	54.568	Gadus_morhua
ENSGALG00000009677	SSB	99	54.279	ENSGAFG00000002749	ssb	99	55.501	Gambusia_affinis
ENSGALG00000009677	SSB	97	55.147	ENSGACG00000005107	ssb	99	55.257	Gasterosteus_aculeatus
ENSGALG00000009677	SSB	100	84.352	ENSGAGG00000007807	SSB	100	85.330	Gopherus_agassizii
ENSGALG00000009677	SSB	99	61.425	ENSGGOG00000024891	-	99	61.425	Gorilla_gorilla
ENSGALG00000009677	SSB	55	70.222	ENSGGOG00000023471	-	92	70.222	Gorilla_gorilla
ENSGALG00000009677	SSB	99	54.726	ENSHBUG00000006584	ssb	99	55.335	Haplochromis_burtoni
ENSGALG00000009677	SSB	100	74.217	ENSHGLG00000017580	-	100	74.217	Heterocephalus_glaber_female
ENSGALG00000009677	SSB	100	70.098	ENSHGLG00000004478	-	100	70.098	Heterocephalus_glaber_female
ENSGALG00000009677	SSB	100	67.157	ENSHGLG00000005832	-	100	67.157	Heterocephalus_glaber_female
ENSGALG00000009677	SSB	100	66.339	ENSHGLG00100008767	-	100	66.339	Heterocephalus_glaber_male
ENSGALG00000009677	SSB	100	74.217	ENSHGLG00100003084	-	100	74.217	Heterocephalus_glaber_male
ENSGALG00000009677	SSB	97	53.865	ENSHCOG00000019011	ssb	99	56.030	Hippocampus_comes
ENSGALG00000009677	SSB	97	59.601	ENSIPUG00000003681	SSB	98	60.199	Ictalurus_punctatus
ENSGALG00000009677	SSB	100	75.184	ENSSTOG00000008416	SSB	100	75.184	Ictidomys_tridecemlineatus
ENSGALG00000009677	SSB	99	64.539	ENSJJAG00000023158	-	99	64.303	Jaculus_jaculus
ENSGALG00000009677	SSB	99	53.827	ENSJJAG00000016250	-	99	53.827	Jaculus_jaculus
ENSGALG00000009677	SSB	99	55.257	ENSKMAG00000017317	ssb	98	56.824	Kryptolebias_marmoratus
ENSGALG00000009677	SSB	99	54.902	ENSLBEG00000017345	ssb	99	56.127	Labrus_bergylta
ENSGALG00000009677	SSB	99	69.024	ENSLACG00000014056	SSB	99	70.443	Latimeria_chalumnae
ENSGALG00000009677	SSB	97	66.667	ENSLOCG00000007933	ssb	98	68.672	Lepisosteus_oculatus
ENSGALG00000009677	SSB	75	59.272	ENSLAFG00000026721	-	99	59.272	Loxodonta_africana
ENSGALG00000009677	SSB	100	70.991	ENSLAFG00000000867	-	100	70.824	Loxodonta_africana
ENSGALG00000009677	SSB	100	63.547	ENSLAFG00000028909	-	100	63.547	Loxodonta_africana
ENSGALG00000009677	SSB	100	74.755	ENSMFAG00000039066	SSB	100	74.755	Macaca_fascicularis
ENSGALG00000009677	SSB	87	66.387	ENSMFAG00000041478	-	100	65.826	Macaca_fascicularis
ENSGALG00000009677	SSB	99	50.985	ENSMMUG00000039889	-	100	50.739	Macaca_mulatta
ENSGALG00000009677	SSB	99	64.339	ENSMMUG00000011724	-	96	63.591	Macaca_mulatta
ENSGALG00000009677	SSB	100	74.755	ENSMMUG00000023051	SSB	100	74.755	Macaca_mulatta
ENSGALG00000009677	SSB	100	64.634	ENSMNEG00000028321	SSB	100	64.634	Macaca_nemestrina
ENSGALG00000009677	SSB	87	66.387	ENSMNEG00000034698	-	100	65.826	Macaca_nemestrina
ENSGALG00000009677	SSB	99	54.115	ENSMLEG00000042644	-	99	53.616	Mandrillus_leucophaeus
ENSGALG00000009677	SSB	100	74.755	ENSMLEG00000035717	SSB	100	74.755	Mandrillus_leucophaeus
ENSGALG00000009677	SSB	99	58.088	ENSMAMG00000004800	ssb	99	59.314	Mastacembelus_armatus
ENSGALG00000009677	SSB	99	54.229	ENSMZEG00005005749	ssb	99	54.839	Maylandia_zebra
ENSGALG00000009677	SSB	100	98.762	ENSMGAG00000010328	SSB	100	98.762	Meleagris_gallopavo
ENSGALG00000009677	SSB	99	75.795	ENSMICG00000028252	-	99	75.795	Microcebus_murinus
ENSGALG00000009677	SSB	98	69.750	ENSMICG00000033304	-	96	72.368	Microcebus_murinus
ENSGALG00000009677	SSB	100	71.292	ENSMODG00000008227	-	100	72.010	Monodelphis_domestica
ENSGALG00000009677	SSB	100	57.108	ENSMODG00000007027	-	100	57.843	Monodelphis_domestica
ENSGALG00000009677	SSB	85	77.987	ENSMODG00000019271	-	99	77.987	Monodelphis_domestica
ENSGALG00000009677	SSB	89	54.400	ENSMALG00000008045	ssb	92	56.667	Monopterus_albus
ENSGALG00000009677	SSB	99	65.238	MGP_CAROLIEiJ_G0023629	-	100	76.630	Mus_caroli
ENSGALG00000009677	SSB	99	64.524	ENSMUSG00000068882	Ssb	100	76.630	Mus_musculus
ENSGALG00000009677	SSB	99	64.200	MGP_PahariEiJ_G0025081	-	100	76.087	Mus_pahari
ENSGALG00000009677	SSB	99	64.524	MGP_SPRETEiJ_G0024551	Ssb	100	76.630	Mus_spretus
ENSGALG00000009677	SSB	100	75.490	ENSMPUG00000000785	SSB	100	75.490	Mustela_putorius_furo
ENSGALG00000009677	SSB	100	72.577	ENSMLUG00000016316	SSB	100	72.577	Myotis_lucifugus
ENSGALG00000009677	SSB	99	54.726	ENSNBRG00000012661	ssb	99	55.335	Neolamprologus_brichardi
ENSGALG00000009677	SSB	100	55.637	ENSNLEG00000028754	-	100	55.637	Nomascus_leucogenys
ENSGALG00000009677	SSB	100	74.755	ENSNLEG00000004958	SSB	100	74.755	Nomascus_leucogenys
ENSGALG00000009677	SSB	61	81.048	ENSMEUG00000001650	-	100	81.048	Notamacropus_eugenii
ENSGALG00000009677	SSB	72	57.439	ENSOPRG00000008475	-	100	59.655	Ochotona_princeps
ENSGALG00000009677	SSB	93	72.679	ENSODEG00000004202	-	99	72.679	Octodon_degus
ENSGALG00000009677	SSB	99	54.975	ENSONIG00000008776	ssb	99	55.583	Oreochromis_niloticus
ENSGALG00000009677	SSB	99	75.743	ENSOANG00000008543	SSB	99	76.485	Ornithorhynchus_anatinus
ENSGALG00000009677	SSB	100	74.146	ENSOCUG00000010396	SSB	100	74.878	Oryctolagus_cuniculus
ENSGALG00000009677	SSB	99	56.127	ENSORLG00000017208	ssb	99	56.968	Oryzias_latipes
ENSGALG00000009677	SSB	99	55.882	ENSORLG00020022574	ssb	99	56.724	Oryzias_latipes_hni
ENSGALG00000009677	SSB	99	55.882	ENSORLG00015016592	ssb	99	57.108	Oryzias_latipes_hsok
ENSGALG00000009677	SSB	99	56.757	ENSOMEG00000003193	ssb	99	59.214	Oryzias_melastigma
ENSGALG00000009677	SSB	100	70.353	ENSOGAG00000029768	SSB	100	71.059	Otolemur_garnettii
ENSGALG00000009677	SSB	100	75.490	ENSOARG00000002906	SSB	100	75.490	Ovis_aries
ENSGALG00000009677	SSB	99	60.688	ENSPPAG00000032427	-	100	68.992	Pan_paniscus
ENSGALG00000009677	SSB	100	74.265	ENSPPAG00000042673	SSB	100	74.265	Pan_paniscus
ENSGALG00000009677	SSB	96	58.098	ENSPPAG00000006189	-	92	60.894	Pan_paniscus
ENSGALG00000009677	SSB	100	74.328	ENSPPRG00000006599	SSB	100	74.328	Panthera_pardus
ENSGALG00000009677	SSB	100	74.328	ENSPTIG00000011021	SSB	97	74.328	Panthera_tigris_altaica
ENSGALG00000009677	SSB	100	74.265	ENSPTRG00000012620	SSB	100	74.265	Pan_troglodytes
ENSGALG00000009677	SSB	99	58.231	ENSPTRG00000042880	-	99	56.511	Pan_troglodytes
ENSGALG00000009677	SSB	100	54.300	ENSPANG00000034126	-	100	54.545	Papio_anubis
ENSGALG00000009677	SSB	100	74.755	ENSPANG00000007440	SSB	100	74.755	Papio_anubis
ENSGALG00000009677	SSB	96	55.670	ENSPANG00000031877	-	92	55.155	Papio_anubis
ENSGALG00000009677	SSB	99	63.145	ENSPKIG00000019325	ssb	99	64.865	Paramormyrops_kingsleyae
ENSGALG00000009677	SSB	100	82.885	ENSPSIG00000002265	SSB	93	83.863	Pelodiscus_sinensis
ENSGALG00000009677	SSB	86	52.825	ENSPMGG00000020075	ssb	92	53.955	Periophthalmus_magnuspinnatus
ENSGALG00000009677	SSB	99	47.086	ENSPMAG00000002982	ssb	100	46.620	Petromyzon_marinus
ENSGALG00000009677	SSB	100	74.083	ENSPCIG00000028591	-	100	75.795	Phascolarctos_cinereus
ENSGALG00000009677	SSB	99	54.768	ENSPFOG00000009650	ssb	99	55.990	Poecilia_formosa
ENSGALG00000009677	SSB	99	54.768	ENSPLAG00000020498	ssb	99	56.235	Poecilia_latipinna
ENSGALG00000009677	SSB	99	54.523	ENSPMEG00000001051	ssb	99	55.610	Poecilia_mexicana
ENSGALG00000009677	SSB	91	57.294	ENSPREG00000003708	ssb	98	58.621	Poecilia_reticulata
ENSGALG00000009677	SSB	100	74.755	ENSPPYG00000012919	SSB	100	74.755	Pongo_abelii
ENSGALG00000009677	SSB	95	70.984	ENSPCAG00000010276	SSB	100	71.762	Procavia_capensis
ENSGALG00000009677	SSB	100	72.683	ENSPCOG00000015741	-	100	71.707	Propithecus_coquereli
ENSGALG00000009677	SSB	100	66.176	ENSPCOG00000023189	-	100	66.176	Propithecus_coquereli
ENSGALG00000009677	SSB	85	78.551	ENSPVAG00000006615	SSB	98	78.551	Pteropus_vampyrus
ENSGALG00000009677	SSB	99	54.229	ENSPNYG00000006444	ssb	99	54.839	Pundamilia_nyererei
ENSGALG00000009677	SSB	97	56.219	ENSPNAG00000011708	ssb	99	57.317	Pygocentrus_nattereri
ENSGALG00000009677	SSB	99	65.356	ENSRNOG00000011606	AABR07061825.1	99	65.356	Rattus_norvegicus
ENSGALG00000009677	SSB	99	65.000	ENSRNOG00000007998	Ssb	99	65.000	Rattus_norvegicus
ENSGALG00000009677	SSB	93	64.021	ENSRBIG00000041232	-	97	64.021	Rhinopithecus_bieti
ENSGALG00000009677	SSB	100	61.017	ENSRBIG00000041972	SSB	88	77.391	Rhinopithecus_bieti
ENSGALG00000009677	SSB	89	69.061	ENSRBIG00000041608	-	98	69.061	Rhinopithecus_bieti
ENSGALG00000009677	SSB	100	52.941	ENSRBIG00000029918	-	100	51.471	Rhinopithecus_bieti
ENSGALG00000009677	SSB	89	71.507	ENSRROG00000034102	-	98	67.332	Rhinopithecus_roxellana
ENSGALG00000009677	SSB	97	64.810	ENSRROG00000031684	-	99	64.810	Rhinopithecus_roxellana
ENSGALG00000009677	SSB	100	74.755	ENSRROG00000040346	SSB	100	74.755	Rhinopithecus_roxellana
ENSGALG00000009677	SSB	100	61.369	ENSRROG00000007149	-	100	61.369	Rhinopithecus_roxellana
ENSGALG00000009677	SSB	100	74.755	ENSSBOG00000027204	SSB	100	74.755	Saimiri_boliviensis_boliviensis
ENSGALG00000009677	SSB	100	73.350	ENSSHAG00000018062	SSB	100	75.795	Sarcophilus_harrisii
ENSGALG00000009677	SSB	99	61.858	ENSSFOG00015000550	ssb	99	64.059	Scleropages_formosus
ENSGALG00000009677	SSB	97	53.571	ENSSMAG00000013885	ssb	99	57.641	Scophthalmus_maximus
ENSGALG00000009677	SSB	99	56.863	ENSSDUG00000020117	ssb	99	58.088	Seriola_dumerili
ENSGALG00000009677	SSB	99	54.568	ENSSLDG00000002525	ssb	99	55.802	Seriola_lalandi_dorsalis
ENSGALG00000009677	SSB	95	67.792	ENSSARG00000007747	SSB	100	67.792	Sorex_araneus
ENSGALG00000009677	SSB	100	80.098	ENSSPUG00000012112	SSB	100	80.835	Sphenodon_punctatus
ENSGALG00000009677	SSB	99	54.902	ENSSPAG00000005696	ssb	99	56.127	Stegastes_partitus
ENSGALG00000009677	SSB	98	65.500	ENSSSCG00000015938	SSB	98	65.500	Sus_scrofa
ENSGALG00000009677	SSB	100	93.366	ENSTGUG00000007942	SSB	100	94.103	Taeniopygia_guttata
ENSGALG00000009677	SSB	99	57.143	ENSTRUG00000017839	ssb	99	58.374	Takifugu_rubripes
ENSGALG00000009677	SSB	90	60.165	ENSTNIG00000000915	ssb	100	60.165	Tetraodon_nigroviridis
ENSGALG00000009677	SSB	82	77.391	ENSTBEG00000009056	SSB	82	77.391	Tupaia_belangeri
ENSGALG00000009677	SSB	100	74.878	ENSTTRG00000002152	SSB	100	74.634	Tursiops_truncatus
ENSGALG00000009677	SSB	90	78.533	ENSUAMG00000027833	SSB	97	78.533	Ursus_americanus
ENSGALG00000009677	SSB	100	75.306	ENSUMAG00000013907	SSB	100	75.306	Ursus_maritimus
ENSGALG00000009677	SSB	100	74.755	ENSVPAG00000007183	SSB	100	74.755	Vicugna_pacos
ENSGALG00000009677	SSB	100	75.061	ENSVVUG00000016573	SSB	78	75.061	Vulpes_vulpes
ENSGALG00000009677	SSB	99	58.920	ENSXETG00000025402	ssb	99	60.471	Xenopus_tropicalis
ENSGALG00000009677	SSB	92	54.474	ENSXCOG00000019109	ssb	93	62.500	Xiphophorus_couchianus
ENSGALG00000009677	SSB	99	54.523	ENSXMAG00000004888	ssb	99	55.746	Xiphophorus_maculatus