EuRBPDB

Description

RBDs

Transcripts

Gene Model

PPI

Paralogs

Orthologs

Description

Ensembl ID: ENSHGLG00000008434 (Gene tree)
Gene ID: -
Gene Symbol: PABPC1L
Alias: C20orf119|PABPC1L1|dJ1069P2.3|ePAB
Full Name: poly(A) binding protein cytoplasmic 1 like
Gene Type: protein_coding
Species: Heterocephalus_glaber_female
Status: putative
Strand: Minus strand
Length: 25728 bases
Position: chrJH602103.1:9545145-9570872
Accession: 101704744
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
ENSHGLP00000011773	PABP	PF00658.18	2e-26	1	1
ENSHGLP00000011773	RRM_5	PF13893.6	1.3e-11	1	1
ENSHGLP00000011773	RRM_1	PF00076.22	3.3e-88	1	4
ENSHGLP00000011773	RRM_1	PF00076.22	3.3e-88	2	4
ENSHGLP00000011773	RRM_1	PF00076.22	3.3e-88	3	4
ENSHGLP00000011773	RRM_1	PF00076.22	3.3e-88	4	4

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
ENSHGLT00000011901	PABPC1L-201	2991	XM_004858177	ENSHGLP00000011773	613 (aa)	XP_004858234	UPI00034F379D

Gene Model

Click here to download ENSHGLG00000008434's gene model file

Protein-Protein Interaction (PPI)

Clik here to download ENSHGLG00000008434's network

Paralogs

Ensembl ID	Gene Symbol	Coverage	Identiy	Paralog	Gene Symbol	Coverage	Identiy
ENSHGLG00000008434	PABPC1L	99	63.265	ENSHGLG00000011162	PABPC4	98	64.412
ENSHGLG00000008434	PABPC1L	60	71.429	ENSHGLG00000019668	-	98	71.429
ENSHGLG00000008434	PABPC1L	99	67.250	ENSHGLG00000007713	-	98	67.250
ENSHGLG00000008434	PABPC1L	61	33.735	ENSHGLG00000005423	ELAVL2	99	33.735
ENSHGLG00000008434	PABPC1L	62	62.259	ENSHGLG00000020104	PABPC4L	98	62.259
ENSHGLG00000008434	PABPC1L	54	36.047	ENSHGLG00000017139	CELF2	66	36.047
ENSHGLG00000008434	PABPC1L	59	35.450	ENSHGLG00000012360	SF3B4	54	35.979
ENSHGLG00000008434	PABPC1L	57	30.323	ENSHGLG00000003139	HNRNPA1	59	30.323
ENSHGLG00000008434	PABPC1L	98	51.792	ENSHGLG00000003042	-	94	51.434
ENSHGLG00000008434	PABPC1L	64	32.000	ENSHGLG00000010899	RBM19	73	32.000
ENSHGLG00000008434	PABPC1L	57	30.675	ENSHGLG00000005451	HNRNPA3	50	30.323
ENSHGLG00000008434	PABPC1L	61	38.889	ENSHGLG00000015199	ELAVL3	92	32.530
ENSHGLG00000008434	PABPC1L	60	71.429	ENSHGLG00000019762	-	98	71.429
ENSHGLG00000008434	PABPC1L	50	35.294	ENSHGLG00000006045	CELF3	76	34.247
ENSHGLG00000008434	PABPC1L	59	30.526	ENSHGLG00000008478	-	90	30.636

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
ENSHGLG00000008434	PABPC1L	100	83.710	ENSG00000101104	PABPC1L	99	90.432	Homo_sapiens
ENSHGLG00000008434	PABPC1L	98	64.412	ENSAPOG00000010842	pabpc1l	98	64.573	Acanthochromis_polyacanthus
ENSHGLG00000008434	PABPC1L	99	74.259	ENSAMEG00000004099	PABPC1L	100	74.259	Ailuropoda_melanoleuca
ENSHGLG00000008434	PABPC1L	99	66.592	ENSACIG00000021004	-	100	70.803	Amphilophus_citrinellus
ENSHGLG00000008434	PABPC1L	99	64.184	ENSAOCG00000006167	-	99	64.172	Amphiprion_ocellaris
ENSHGLG00000008434	PABPC1L	99	68.360	ENSAPEG00000010558	-	97	77.228	Amphiprion_percula
ENSHGLG00000008434	PABPC1L	99	65.971	ENSATEG00000007428	pabpc1l	100	65.971	Anabas_testudineus
ENSHGLG00000008434	PABPC1L	98	71.637	ENSAPLG00000008415	PABPC1L	98	71.729	Anas_platyrhynchos
ENSHGLG00000008434	PABPC1L	99	71.040	ENSACAG00000011841	PABPC1L	98	71.040	Anolis_carolinensis
ENSHGLG00000008434	PABPC1L	100	81.452	ENSANAG00000026826	PABPC1L	100	81.452	Aotus_nancymaae
ENSHGLG00000008434	PABPC1L	99	64.771	ENSACLG00000024791	pabpc1l	100	64.455	Astatotilapia_calliptera
ENSHGLG00000008434	PABPC1L	99	67.038	ENSAMXG00000013552	pabpc1l	99	66.561	Astyanax_mexicanus
ENSHGLG00000008434	PABPC1L	55	93.452	ENSBTAG00000055300	-	100	93.452	Bos_taurus
ENSHGLG00000008434	PABPC1L	67	73.852	ENSBTAG00000016849	PABPC1L	84	73.852	Bos_taurus
ENSHGLG00000008434	PABPC1L	100	81.935	ENSCJAG00000017628	PABPC1L	100	81.935	Callithrix_jacchus
ENSHGLG00000008434	PABPC1L	60	74.713	ENSCJAG00000038506	-	100	74.713	Callithrix_jacchus
ENSHGLG00000008434	PABPC1L	99	73.733	ENSCAFG00000009573	PABPC1L	100	73.190	Canis_familiaris
ENSHGLG00000008434	PABPC1L	99	82.602	ENSCAFG00020013828	PABPC1L	99	82.602	Canis_lupus_dingo
ENSHGLG00000008434	PABPC1L	97	80.528	ENSCHIG00000025759	PABPC1L	91	80.528	Capra_hircus
ENSHGLG00000008434	PABPC1L	100	80.806	ENSTSYG00000006645	PABPC1L	100	80.806	Carlito_syrichta
ENSHGLG00000008434	PABPC1L	99	89.617	ENSCAPG00000014519	PABPC1L	100	89.617	Cavia_aperea
ENSHGLG00000008434	PABPC1L	100	93.215	ENSCPOG00000013633	PABPC1L	100	93.215	Cavia_porcellus
ENSHGLG00000008434	PABPC1L	100	81.935	ENSCCAG00000023338	PABPC1L	100	81.935	Cebus_capucinus
ENSHGLG00000008434	PABPC1L	100	83.548	ENSCATG00000034065	PABPC1L	100	83.548	Cercocebus_atys
ENSHGLG00000008434	PABPC1L	100	92.892	ENSCLAG00000005298	PABPC1L	100	92.892	Chinchilla_lanigera
ENSHGLG00000008434	PABPC1L	100	83.710	ENSCSAG00000014242	PABPC1L	100	83.710	Chlorocebus_sabaeus
ENSHGLG00000008434	PABPC1L	72	79.638	ENSCPBG00000017755	-	99	79.638	Chrysemys_picta_bellii
ENSHGLG00000008434	PABPC1L	53	78.788	ENSCPBG00000007810	-	83	78.788	Chrysemys_picta_bellii
ENSHGLG00000008434	PABPC1L	97	48.497	ENSCING00000002796	-	97	50.000	Ciona_intestinalis
ENSHGLG00000008434	PABPC1L	97	59.024	ENSCSAVG00000004246	-	86	79.518	Ciona_savignyi
ENSHGLG00000008434	PABPC1L	97	56.073	ENSCSAVG00000003102	-	100	78.767	Ciona_savignyi
ENSHGLG00000008434	PABPC1L	100	79.173	ENSCANG00000037530	PABPC1L	100	79.173	Colobus_angolensis_palliatus
ENSHGLG00000008434	PABPC1L	99	77.273	ENSCGRG00001021938	Pabpc1l	99	77.273	Cricetulus_griseus_chok1gshd
ENSHGLG00000008434	PABPC1L	99	77.273	ENSCGRG00000007946	Pabpc1l	99	77.273	Cricetulus_griseus_crigri
ENSHGLG00000008434	PABPC1L	99	63.201	ENSCSEG00000021295	pabpc1l	99	63.036	Cynoglossus_semilaevis
ENSHGLG00000008434	PABPC1L	99	67.261	ENSCVAG00000015943	-	86	81.679	Cyprinodon_variegatus
ENSHGLG00000008434	PABPC1L	99	67.943	ENSDARG00000002597	pabpc1l	88	83.846	Danio_rerio
ENSHGLG00000008434	PABPC1L	99	83.683	ENSDNOG00000048450	PABPC1L	99	83.683	Dasypus_novemcinctus
ENSHGLG00000008434	PABPC1L	99	83.495	ENSDORG00000001067	Pabpc1l	99	83.495	Dipodomys_ordii
ENSHGLG00000008434	PABPC1L	96	86.585	ENSETEG00000009969	PABPC1L	73	86.585	Echinops_telfairi
ENSHGLG00000008434	PABPC1L	53	84.615	ENSEASG00005023407	-	73	81.111	Equus_asinus_asinus
ENSHGLG00000008434	PABPC1L	56	93.452	ENSEASG00005023408	-	100	93.452	Equus_asinus_asinus
ENSHGLG00000008434	PABPC1L	68	80.080	ENSECAG00000040139	-	89	80.080	Equus_caballus
ENSHGLG00000008434	PABPC1L	100	79.160	ENSEEUG00000005978	PABPC1L	100	79.160	Erinaceus_europaeus
ENSHGLG00000008434	PABPC1L	99	68.189	ENSELUG00000001811	pabpc1l	100	67.874	Esox_lucius
ENSHGLG00000008434	PABPC1L	99	83.415	ENSFCAG00000011692	PABPC1L	99	83.577	Felis_catus
ENSHGLG00000008434	PABPC1L	83	75.099	ENSFALG00000005372	PABPC1L	85	75.099	Ficedula_albicollis
ENSHGLG00000008434	PABPC1L	100	92.170	ENSFDAG00000008291	PABPC1L	100	91.517	Fukomys_damarensis
ENSHGLG00000008434	PABPC1L	99	66.451	ENSFHEG00000003398	pabpc1l	99	66.126	Fundulus_heteroclitus
ENSHGLG00000008434	PABPC1L	99	66.990	ENSGMOG00000004437	pabpc1l	98	66.990	Gadus_morhua
ENSHGLG00000008434	PABPC1L	98	68.503	ENSGALG00000004123	PABPC1L	98	82.022	Gallus_gallus
ENSHGLG00000008434	PABPC1L	99	62.787	ENSGAFG00000021057	pabpc1l	99	62.481	Gambusia_affinis
ENSHGLG00000008434	PABPC1L	99	64.320	ENSGACG00000002792	pabpc1l	99	64.160	Gasterosteus_aculeatus
ENSHGLG00000008434	PABPC1L	99	73.046	ENSGAGG00000000589	PABPC1L	96	82.955	Gopherus_agassizii
ENSHGLG00000008434	PABPC1L	100	83.871	ENSGGOG00000004584	PABPC1L	100	83.871	Gorilla_gorilla
ENSHGLG00000008434	PABPC1L	99	64.613	ENSHBUG00000006134	pabpc1l	100	64.297	Haplochromis_burtoni
ENSHGLG00000008434	PABPC1L	100	100.000	ENSHGLG00100012151	PABPC1L	100	100.000	Heterocephalus_glaber_male
ENSHGLG00000008434	PABPC1L	60	79.213	ENSHGLG00100019506	-	100	79.213	Heterocephalus_glaber_male
ENSHGLG00000008434	PABPC1L	99	64.073	ENSHCOG00000009270	pabpc1l	99	64.833	Hippocampus_comes
ENSHGLG00000008434	PABPC1L	60	59.770	ENSIPUG00000006780	-	85	59.770	Ictalurus_punctatus
ENSHGLG00000008434	PABPC1L	99	67.512	ENSIPUG00000012602	pabpc1l	99	67.195	Ictalurus_punctatus
ENSHGLG00000008434	PABPC1L	60	59.195	ENSIPUG00000006760	-	88	59.195	Ictalurus_punctatus
ENSHGLG00000008434	PABPC1L	100	85.000	ENSSTOG00000003739	PABPC1L	100	85.000	Ictidomys_tridecemlineatus
ENSHGLG00000008434	PABPC1L	99	70.513	ENSJJAG00000022406	Pabpc1l	99	68.137	Jaculus_jaculus
ENSHGLG00000008434	PABPC1L	99	65.811	ENSKMAG00000016847	pabpc1l	100	65.329	Kryptolebias_marmoratus
ENSHGLG00000008434	PABPC1L	99	63.191	ENSLBEG00000010084	pabpc1l	100	63.349	Labrus_bergylta
ENSHGLG00000008434	PABPC1L	99	66.667	ENSLACG00000005762	PABPC1L	98	66.193	Latimeria_chalumnae
ENSHGLG00000008434	PABPC1L	99	68.530	ENSLOCG00000007519	pabpc1l	98	68.581	Lepisosteus_oculatus
ENSHGLG00000008434	PABPC1L	99	80.990	ENSLAFG00000003553	PABPC1L	100	80.990	Loxodonta_africana
ENSHGLG00000008434	PABPC1L	100	83.387	ENSMFAG00000040365	PABPC1L	100	83.387	Macaca_fascicularis
ENSHGLG00000008434	PABPC1L	100	81.059	ENSMNEG00000032298	PABPC1L	100	81.059	Macaca_nemestrina
ENSHGLG00000008434	PABPC1L	100	83.548	ENSMLEG00000043139	PABPC1L	100	83.548	Mandrillus_leucophaeus
ENSHGLG00000008434	PABPC1L	97	64.851	ENSMAMG00000011854	pabpc1l	97	64.851	Mastacembelus_armatus
ENSHGLG00000008434	PABPC1L	99	64.771	ENSMZEG00005023713	pabpc1l	100	64.455	Maylandia_zebra
ENSHGLG00000008434	PABPC1L	99	73.151	ENSMGAG00000003674	PABPC1L	98	73.151	Meleagris_gallopavo
ENSHGLG00000008434	PABPC1L	99	75.980	ENSMOCG00000013186	Pabpc1l	99	75.980	Microtus_ochrogaster
ENSHGLG00000008434	PABPC1L	99	65.751	ENSMMOG00000020574	-	90	82.308	Mola_mola
ENSHGLG00000008434	PABPC1L	94	75.298	ENSMODG00000016198	PABPC1L	99	75.331	Monodelphis_domestica
ENSHGLG00000008434	PABPC1L	97	63.339	ENSMALG00000021622	pabpc1l	100	63.339	Monopterus_albus
ENSHGLG00000008434	PABPC1L	99	76.672	MGP_CAROLIEiJ_G0024627	Pabpc1l	99	76.672	Mus_caroli
ENSHGLG00000008434	PABPC1L	99	76.346	ENSMUSG00000054582	Pabpc1l	99	76.346	Mus_musculus
ENSHGLG00000008434	PABPC1L	99	76.710	MGP_PahariEiJ_G0026071	Pabpc1l	99	76.710	Mus_pahari
ENSHGLG00000008434	PABPC1L	99	76.509	MGP_SPRETEiJ_G0025549	Pabpc1l	99	76.509	Mus_spretus
ENSHGLG00000008434	PABPC1L	99	81.190	ENSMPUG00000017646	PABPC1L	99	81.190	Mustela_putorius_furo
ENSHGLG00000008434	PABPC1L	98	76.329	ENSMLUG00000006324	PABPC1L	97	76.329	Myotis_lucifugus
ENSHGLG00000008434	PABPC1L	99	80.097	ENSNGAG00000010458	Pabpc1l	99	80.097	Nannospalax_galili
ENSHGLG00000008434	PABPC1L	99	64.809	ENSNBRG00000016633	pabpc1l	100	71.277	Neolamprologus_brichardi
ENSHGLG00000008434	PABPC1L	100	65.161	ENSMEUG00000007905	-	100	65.161	Notamacropus_eugenii
ENSHGLG00000008434	PABPC1L	100	81.159	ENSOPRG00000011846	PABPC1L	100	81.159	Ochotona_princeps
ENSHGLG00000008434	PABPC1L	57	75.163	ENSODEG00000002920	-	92	75.163	Octodon_degus
ENSHGLG00000008434	PABPC1L	100	81.301	ENSODEG00000015482	PABPC1L	100	81.301	Octodon_degus
ENSHGLG00000008434	PABPC1L	99	64.929	ENSONIG00000017314	pabpc1l	100	64.613	Oreochromis_niloticus
ENSHGLG00000008434	PABPC1L	89	65.116	ENSOANG00000015793	-	100	65.116	Ornithorhynchus_anatinus
ENSHGLG00000008434	PABPC1L	99	75.382	ENSOCUG00000003005	PABPC1L	100	75.382	Oryctolagus_cuniculus
ENSHGLG00000008434	PABPC1L	99	69.930	ENSORLG00000010322	pabpc1l	100	72.941	Oryzias_latipes
ENSHGLG00000008434	PABPC1L	99	69.930	ENSORLG00020021193	pabpc1l	100	72.941	Oryzias_latipes_hni
ENSHGLG00000008434	PABPC1L	99	69.930	ENSORLG00015017018	pabpc1l	100	71.765	Oryzias_latipes_hsok
ENSHGLG00000008434	PABPC1L	99	61.480	ENSOMEG00000010789	pabpc1l	100	61.178	Oryzias_melastigma
ENSHGLG00000008434	PABPC1L	99	80.794	ENSOGAG00000032999	PABPC1L	100	80.794	Otolemur_garnettii
ENSHGLG00000008434	PABPC1L	99	81.818	ENSOARG00000004578	PABPC1L	99	81.818	Ovis_aries
ENSHGLG00000008434	PABPC1L	100	83.548	ENSPPAG00000039677	PABPC1L	100	83.548	Pan_paniscus
ENSHGLG00000008434	PABPC1L	63	78.395	ENSPPRG00000019327	-	98	78.395	Panthera_pardus
ENSHGLG00000008434	PABPC1L	99	83.744	ENSPTIG00000006524	PABPC1L	99	83.908	Panthera_tigris_altaica
ENSHGLG00000008434	PABPC1L	63	90.123	ENSPTRG00000013531	PABPC1L	99	90.123	Pan_troglodytes
ENSHGLG00000008434	PABPC1L	100	83.548	ENSPANG00000018788	PABPC1L	100	83.548	Papio_anubis
ENSHGLG00000008434	PABPC1L	99	68.000	ENSPKIG00000007322	pabpc1l	99	67.840	Paramormyrops_kingsleyae
ENSHGLG00000008434	PABPC1L	53	79.231	ENSPKIG00000016810	-	93	79.231	Paramormyrops_kingsleyae
ENSHGLG00000008434	PABPC1L	68	79.856	ENSPSIG00000013289	PABPC1L	99	79.856	Pelodiscus_sinensis
ENSHGLG00000008434	PABPC1L	72	64.141	ENSPMGG00000023121	-	92	64.141	Periophthalmus_magnuspinnatus
ENSHGLG00000008434	PABPC1L	61	74.419	ENSPMGG00000023117	pabpc1l	92	81.061	Periophthalmus_magnuspinnatus
ENSHGLG00000008434	PABPC1L	99	76.672	ENSPEMG00000020671	Pabpc1l	99	76.672	Peromyscus_maniculatus_bairdii
ENSHGLG00000008434	PABPC1L	99	67.541	ENSPFOG00000013241	pabpc1l	99	66.992	Poecilia_formosa
ENSHGLG00000008434	PABPC1L	99	66.184	ENSPLAG00000017490	-	100	65.655	Poecilia_latipinna
ENSHGLG00000008434	PABPC1L	99	66.559	ENSPMEG00000007677	pabpc1l	99	66.236	Poecilia_mexicana
ENSHGLG00000008434	PABPC1L	66	74.877	ENSPREG00000015886	pabpc1l	99	74.877	Poecilia_reticulata
ENSHGLG00000008434	PABPC1L	100	83.871	ENSPPYG00000011042	PABPC1L	100	83.871	Pongo_abelii
ENSHGLG00000008434	PABPC1L	62	85.938	ENSPCAG00000011787	-	75	85.938	Procavia_capensis
ENSHGLG00000008434	PABPC1L	99	82.871	ENSPCOG00000020630	PABPC1L	99	82.871	Propithecus_coquereli
ENSHGLG00000008434	PABPC1L	99	80.195	ENSPVAG00000012663	PABPC1L	99	80.195	Pteropus_vampyrus
ENSHGLG00000008434	PABPC1L	99	63.961	ENSPNYG00000004101	pabpc1l	100	71.277	Pundamilia_nyererei
ENSHGLG00000008434	PABPC1L	99	61.404	ENSPNAG00000021248	pabpc1l	99	61.085	Pygocentrus_nattereri
ENSHGLG00000008434	PABPC1L	99	76.375	ENSRNOG00000012750	Pabpc1l	100	76.375	Rattus_norvegicus
ENSHGLG00000008434	PABPC1L	100	83.710	ENSRBIG00000035381	PABPC1L	100	83.710	Rhinopithecus_bieti
ENSHGLG00000008434	PABPC1L	100	83.387	ENSRROG00000044498	PABPC1L	100	83.387	Rhinopithecus_roxellana
ENSHGLG00000008434	PABPC1L	100	81.290	ENSSBOG00000009532	-	100	81.290	Saimiri_boliviensis_boliviensis
ENSHGLG00000008434	PABPC1L	99	69.838	ENSSHAG00000011446	PABPC1L	100	69.838	Sarcophilus_harrisii
ENSHGLG00000008434	PABPC1L	99	68.949	ENSSFOG00015017304	pabpc1l	99	68.790	Scleropages_formosus
ENSHGLG00000008434	PABPC1L	99	63.854	ENSSMAG00000010516	pabpc1l	98	82.171	Scophthalmus_maximus
ENSHGLG00000008434	PABPC1L	99	64.455	ENSSDUG00000014743	pabpc1l	100	64.455	Seriola_dumerili
ENSHGLG00000008434	PABPC1L	99	71.223	ENSSLDG00000006981	pabpc1l	95	76.555	Seriola_lalandi_dorsalis
ENSHGLG00000008434	PABPC1L	99	70.335	ENSSPUG00000004233	PABPC1L	98	70.335	Sphenodon_punctatus
ENSHGLG00000008434	PABPC1L	99	63.576	ENSSPAG00000017137	pabpc1l	99	63.742	Stegastes_partitus
ENSHGLG00000008434	PABPC1L	99	80.619	ENSSSCG00000007382	PABPC1L	99	80.619	Sus_scrofa
ENSHGLG00000008434	PABPC1L	99	70.295	ENSTGUG00000005114	PABPC1L	99	70.295	Taeniopygia_guttata
ENSHGLG00000008434	PABPC1L	97	65.253	ENSTRUG00000011085	pabpc1l	97	65.253	Takifugu_rubripes
ENSHGLG00000008434	PABPC1L	100	62.047	ENSTNIG00000017903	pabpc1l	99	62.047	Tetraodon_nigroviridis
ENSHGLG00000008434	PABPC1L	94	89.726	ENSTBEG00000002851	PABPC1L	80	89.726	Tupaia_belangeri
ENSHGLG00000008434	PABPC1L	94	79.897	ENSTTRG00000007618	PABPC1L	100	80.100	Tursiops_truncatus
ENSHGLG00000008434	PABPC1L	99	82.178	ENSUAMG00000017682	PABPC1L	99	82.178	Ursus_americanus
ENSHGLG00000008434	PABPC1L	69	87.534	ENSUMAG00000025436	PABPC1L	94	87.534	Ursus_maritimus
ENSHGLG00000008434	PABPC1L	99	89.723	ENSVVUG00000018939	PABPC1L	99	89.723	Vulpes_vulpes
ENSHGLG00000008434	PABPC1L	99	66.987	ENSXETG00000022831	pabpc1l	98	66.827	Xenopus_tropicalis
ENSHGLG00000008434	PABPC1L	99	69.811	ENSXCOG00000005459	pabpc1l	99	78.723	Xiphophorus_couchianus
ENSHGLG00000008434	PABPC1L	99	66.023	ENSXMAG00000017135	pabpc1l	100	65.495	Xiphophorus_maculatus