EuRBPDB

Description

RBDs

Transcripts

Gene Model

Pathways

PPI

Paralogs

Orthologs

Description

Ensembl ID: ENSMALG00000005885 (Gene tree)
Gene ID: 109961947
Gene Symbol: eif4ebp2
Alias: fa55e11|wu:fa55e11|wu:fp45e09
Full Name: eukaryotic translation initiation factor 4E binding protein 2
Gene Type: protein_coding
Species: Monopterus_albus
Status: putative
Strand: Plus strand
Length: 6130 bases
Position: chrKV884748.1:2611628-2617757
Accession: 109961947
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
ENSMALP00000008270	eIF_4EBP	PF05456.11	1.6e-47	1	1

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
ENSMALT00000008446	-	3321	XM_020603222	ENSMALP00000008270	114 (aa)	XP_020458878	UPI0009B36796

Gene Model

Click here to download ENSMALG00000005885's gene model file

Pathways

Pathway ID	Pathway Name	Source
malb03013	RNA transport	KEGG

Protein-Protein Interaction (PPI)

Clik here to download ENSMALG00000005885's network

Paralogs

Ensembl ID	Gene Symbol	Coverage	Identiy	Paralog	Gene Symbol	Coverage	Identiy
ENSMALG00000005885	eif4ebp2	100	54.310	ENSMALG00000006261	eif4ebp3l	100	54.386
ENSMALG00000005885	eif4ebp2	100	55.085	ENSMALG00000012932	EIF4EBP1	100	55.085
ENSMALG00000005885	eif4ebp2	100	51.240	ENSMALG00000012228	eif4ebp1	100	51.240

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
ENSMALG00000005885	eif4ebp2	98	73.684	ENSG00000148730	EIF4EBP2	95	74.561	Homo_sapiens
ENSMALG00000005885	eif4ebp2	98	91.071	ENSAPOG00000017183	eif4ebp2	92	91.071	Acanthochromis_polyacanthus
ENSMALG00000005885	eif4ebp2	96	74.775	ENSAMEG00000017848	EIF4EBP2	99	75.676	Ailuropoda_melanoleuca
ENSMALG00000005885	eif4ebp2	100	93.220	ENSACIG00000019336	eif4ebp2	100	93.220	Amphilophus_citrinellus
ENSMALG00000005885	eif4ebp2	100	96.491	ENSAOCG00000022114	eif4ebp2	100	96.491	Amphiprion_ocellaris
ENSMALG00000005885	eif4ebp2	100	96.491	ENSAPEG00000011456	eif4ebp2	100	96.491	Amphiprion_percula
ENSMALG00000005885	eif4ebp2	100	94.737	ENSATEG00000003554	eif4ebp2	100	94.737	Anabas_testudineus
ENSMALG00000005885	eif4ebp2	94	78.505	ENSACAG00000008388	EIF4EBP2	85	78.505	Anolis_carolinensis
ENSMALG00000005885	eif4ebp2	98	74.561	ENSANAG00000024340	EIF4EBP2	95	75.439	Aotus_nancymaae
ENSMALG00000005885	eif4ebp2	100	92.982	ENSACLG00000013067	eif4ebp2	100	92.982	Astatotilapia_calliptera
ENSMALG00000005885	eif4ebp2	100	87.719	ENSAMXG00000037221	eif4ebp2	100	87.719	Astyanax_mexicanus
ENSMALG00000005885	eif4ebp2	98	73.684	ENSBTAG00000053726	EIF4EBP2	95	74.561	Bos_taurus
ENSMALG00000005885	eif4ebp2	98	73.684	ENSCJAG00000040001	EIF4EBP2	95	74.561	Callithrix_jacchus
ENSMALG00000005885	eif4ebp2	98	73.684	ENSCAFG00000014050	EIF4EBP2	95	74.561	Canis_familiaris
ENSMALG00000005885	eif4ebp2	98	73.684	ENSCAFG00020025874	EIF4EBP2	95	74.561	Canis_lupus_dingo
ENSMALG00000005885	eif4ebp2	98	72.807	ENSCHIG00000000417	EIF4EBP2	95	73.684	Capra_hircus
ENSMALG00000005885	eif4ebp2	98	73.684	ENSTSYG00000037472	-	95	74.561	Carlito_syrichta
ENSMALG00000005885	eif4ebp2	98	71.053	ENSTSYG00000036771	-	95	71.930	Carlito_syrichta
ENSMALG00000005885	eif4ebp2	94	74.312	ENSCPOG00000039594	EIF4EBP2	90	75.229	Cavia_porcellus
ENSMALG00000005885	eif4ebp2	98	74.561	ENSCCAG00000037637	EIF4EBP2	95	75.439	Cebus_capucinus
ENSMALG00000005885	eif4ebp2	96	58.559	ENSCATG00000025087	-	97	59.459	Cercocebus_atys
ENSMALG00000005885	eif4ebp2	98	73.684	ENSCATG00000032356	EIF4EBP2	95	74.561	Cercocebus_atys
ENSMALG00000005885	eif4ebp2	98	71.930	ENSCLAG00000013838	EIF4EBP2	95	72.807	Chinchilla_lanigera
ENSMALG00000005885	eif4ebp2	98	73.684	ENSCSAG00000008294	EIF4EBP2	95	74.561	Chlorocebus_sabaeus
ENSMALG00000005885	eif4ebp2	99	76.106	ENSCPBG00000025890	EIF4EBP2	97	76.106	Chrysemys_picta_bellii
ENSMALG00000005885	eif4ebp2	95	62.727	ENSCANG00000017895	-	98	63.636	Colobus_angolensis_palliatus
ENSMALG00000005885	eif4ebp2	61	70.423	ENSCANG00000035781	EIF4EBP2	100	71.831	Colobus_angolensis_palliatus
ENSMALG00000005885	eif4ebp2	96	73.874	ENSCGRG00001010892	Eif4ebp2	93	74.775	Cricetulus_griseus_chok1gshd
ENSMALG00000005885	eif4ebp2	83	73.196	ENSCGRG00000013786	Eif4ebp2	100	74.227	Cricetulus_griseus_crigri
ENSMALG00000005885	eif4ebp2	100	93.860	ENSCSEG00000004926	eif4ebp2	100	93.860	Cynoglossus_semilaevis
ENSMALG00000005885	eif4ebp2	100	91.228	ENSCVAG00000015182	eif4ebp2	100	91.228	Cyprinodon_variegatus
ENSMALG00000005885	eif4ebp2	100	85.965	ENSDARG00000031819	eif4ebp2	100	85.965	Danio_rerio
ENSMALG00000005885	eif4ebp2	98	73.684	ENSDNOG00000010090	EIF4EBP2	95	74.561	Dasypus_novemcinctus
ENSMALG00000005885	eif4ebp2	98	74.561	ENSDORG00000025031	Eif4ebp2	95	75.439	Dipodomys_ordii
ENSMALG00000005885	eif4ebp2	96	58.716	ENSETEG00000013860	EIF4EBP2	92	58.716	Echinops_telfairi
ENSMALG00000005885	eif4ebp2	98	74.561	ENSEASG00005016053	EIF4EBP2	95	75.439	Equus_asinus_asinus
ENSMALG00000005885	eif4ebp2	96	76.577	ENSECAG00000034245	EIF4EBP2	61	75.439	Equus_caballus
ENSMALG00000005885	eif4ebp2	98	73.684	ENSEEUG00000011808	EIF4EBP2	95	74.561	Erinaceus_europaeus
ENSMALG00000005885	eif4ebp2	99	86.957	ENSELUG00000024297	eif4ebp2	97	86.957	Esox_lucius
ENSMALG00000005885	eif4ebp2	98	73.684	ENSFCAG00000036699	EIF4EBP2	95	74.561	Felis_catus
ENSMALG00000005885	eif4ebp2	96	71.818	ENSFALG00000012606	EIF4EBP2	94	71.818	Ficedula_albicollis
ENSMALG00000005885	eif4ebp2	99	86.842	ENSGMOG00000012592	eif4ebp2	97	86.842	Gadus_morhua
ENSMALG00000005885	eif4ebp2	96	64.545	ENSGALG00000037904	EIF4EBP2	95	64.545	Gallus_gallus
ENSMALG00000005885	eif4ebp2	100	94.737	ENSGAFG00000009724	eif4ebp2	100	94.737	Gambusia_affinis
ENSMALG00000005885	eif4ebp2	100	91.304	ENSGACG00000008729	eif4ebp2	97	91.304	Gasterosteus_aculeatus
ENSMALG00000005885	eif4ebp2	99	76.106	ENSGAGG00000017974	EIF4EBP2	97	76.106	Gopherus_agassizii
ENSMALG00000005885	eif4ebp2	98	73.684	ENSGGOG00000036271	EIF4EBP2	95	74.561	Gorilla_gorilla
ENSMALG00000005885	eif4ebp2	96	56.881	ENSGGOG00000041553	-	94	55.357	Gorilla_gorilla
ENSMALG00000005885	eif4ebp2	100	93.860	ENSHBUG00000021120	eif4ebp2	100	93.860	Haplochromis_burtoni
ENSMALG00000005885	eif4ebp2	98	72.807	ENSHGLG00000011838	EIF4EBP2	95	73.684	Heterocephalus_glaber_female
ENSMALG00000005885	eif4ebp2	98	58.261	ENSHGLG00000007653	-	95	58.261	Heterocephalus_glaber_female
ENSMALG00000005885	eif4ebp2	96	59.483	ENSHGLG00000020548	-	93	61.947	Heterocephalus_glaber_female
ENSMALG00000005885	eif4ebp2	99	68.966	ENSHGLG00000009800	-	97	69.828	Heterocephalus_glaber_female
ENSMALG00000005885	eif4ebp2	79	76.667	ENSHGLG00100008743	-	83	74.510	Heterocephalus_glaber_male
ENSMALG00000005885	eif4ebp2	96	56.250	ENSHGLG00100016402	-	92	56.250	Heterocephalus_glaber_male
ENSMALG00000005885	eif4ebp2	96	57.759	ENSHGLG00100000583	-	98	60.169	Heterocephalus_glaber_male
ENSMALG00000005885	eif4ebp2	99	68.966	ENSHGLG00100006847	-	97	69.828	Heterocephalus_glaber_male
ENSMALG00000005885	eif4ebp2	100	78.070	ENSHCOG00000000576	eif4ebp2	98	80.702	Hippocampus_comes
ENSMALG00000005885	eif4ebp2	100	89.474	ENSIPUG00000024675	eif4ebp2	100	89.474	Ictalurus_punctatus
ENSMALG00000005885	eif4ebp2	98	74.561	ENSSTOG00000012937	EIF4EBP2	95	75.439	Ictidomys_tridecemlineatus
ENSMALG00000005885	eif4ebp2	98	75.893	ENSJJAG00000017843	Eif4ebp2	95	75.893	Jaculus_jaculus
ENSMALG00000005885	eif4ebp2	100	95.614	ENSKMAG00000018101	eif4ebp2	100	95.614	Kryptolebias_marmoratus
ENSMALG00000005885	eif4ebp2	100	92.982	ENSLBEG00000001180	eif4ebp2	100	92.982	Labrus_bergylta
ENSMALG00000005885	eif4ebp2	100	78.070	ENSLACG00000016130	EIF4EBP2	100	78.070	Latimeria_chalumnae
ENSMALG00000005885	eif4ebp2	100	78.400	ENSLOCG00000005123	eif4ebp2	99	78.400	Lepisosteus_oculatus
ENSMALG00000005885	eif4ebp2	61	72.222	ENSLAFG00000006809	EIF4EBP2	100	73.611	Loxodonta_africana
ENSMALG00000005885	eif4ebp2	96	58.559	ENSMFAG00000002192	-	97	59.459	Macaca_fascicularis
ENSMALG00000005885	eif4ebp2	100	61.864	ENSMFAG00000000272	-	91	61.864	Macaca_fascicularis
ENSMALG00000005885	eif4ebp2	98	73.684	ENSMFAG00000000143	EIF4EBP2	95	74.561	Macaca_fascicularis
ENSMALG00000005885	eif4ebp2	98	61.207	ENSMMUG00000045244	-	99	61.207	Macaca_mulatta
ENSMALG00000005885	eif4ebp2	98	73.684	ENSMMUG00000023169	EIF4EBP2	95	74.561	Macaca_mulatta
ENSMALG00000005885	eif4ebp2	98	73.684	ENSMNEG00000032389	EIF4EBP2	95	74.561	Macaca_nemestrina
ENSMALG00000005885	eif4ebp2	96	59.459	ENSMNEG00000033973	-	97	60.360	Macaca_nemestrina
ENSMALG00000005885	eif4ebp2	96	61.261	ENSMLEG00000035598	-	99	61.607	Mandrillus_leucophaeus
ENSMALG00000005885	eif4ebp2	61	70.423	ENSMLEG00000034679	EIF4EBP2	100	71.831	Mandrillus_leucophaeus
ENSMALG00000005885	eif4ebp2	100	96.491	ENSMAMG00000021311	eif4ebp2	100	96.491	Mastacembelus_armatus
ENSMALG00000005885	eif4ebp2	100	93.860	ENSMZEG00005027604	eif4ebp2	100	93.860	Maylandia_zebra
ENSMALG00000005885	eif4ebp2	96	73.874	ENSMAUG00000012220	Eif4ebp2	93	74.775	Mesocricetus_auratus
ENSMALG00000005885	eif4ebp2	98	71.053	ENSMICG00000031264	-	95	71.930	Microcebus_murinus
ENSMALG00000005885	eif4ebp2	98	73.684	ENSMICG00000043210	-	95	74.561	Microcebus_murinus
ENSMALG00000005885	eif4ebp2	96	73.874	ENSMOCG00000013203	Eif4ebp2	93	74.775	Microtus_ochrogaster
ENSMALG00000005885	eif4ebp2	100	93.043	ENSMMOG00000010358	eif4ebp2	100	93.043	Mola_mola
ENSMALG00000005885	eif4ebp2	96	79.817	ENSMODG00000012161	EIF4EBP2	96	79.817	Monodelphis_domestica
ENSMALG00000005885	eif4ebp2	97	73.451	MGP_CAROLIEiJ_G0015367	Eif4ebp2	94	74.336	Mus_caroli
ENSMALG00000005885	eif4ebp2	97	73.451	ENSMUSG00000020091	Eif4ebp2	94	74.336	Mus_musculus
ENSMALG00000005885	eif4ebp2	96	73.874	MGP_PahariEiJ_G0030783	Eif4ebp2	93	74.775	Mus_pahari
ENSMALG00000005885	eif4ebp2	97	73.451	MGP_SPRETEiJ_G0016178	Eif4ebp2	94	74.336	Mus_spretus
ENSMALG00000005885	eif4ebp2	98	73.684	ENSMPUG00000004300	EIF4EBP2	95	74.561	Mustela_putorius_furo
ENSMALG00000005885	eif4ebp2	61	70.833	ENSMLUG00000015295	EIF4EBP2	100	72.222	Myotis_lucifugus
ENSMALG00000005885	eif4ebp2	96	74.775	ENSNGAG00000021168	Eif4ebp2	93	75.676	Nannospalax_galili
ENSMALG00000005885	eif4ebp2	100	93.860	ENSNBRG00000018719	eif4ebp2	100	93.860	Neolamprologus_brichardi
ENSMALG00000005885	eif4ebp2	98	74.561	ENSNLEG00000016121	-	95	75.439	Nomascus_leucogenys
ENSMALG00000005885	eif4ebp2	94	62.385	ENSNLEG00000027712	-	100	63.303	Nomascus_leucogenys
ENSMALG00000005885	eif4ebp2	96	54.955	ENSNLEG00000029057	-	94	54.386	Nomascus_leucogenys
ENSMALG00000005885	eif4ebp2	61	75.362	ENSMEUG00000011831	EIF4EBP2	100	75.362	Notamacropus_eugenii
ENSMALG00000005885	eif4ebp2	98	73.684	ENSOPRG00000015819	EIF4EBP2	95	74.561	Ochotona_princeps
ENSMALG00000005885	eif4ebp2	98	75.000	ENSODEG00000014467	EIF4EBP2	95	75.000	Octodon_degus
ENSMALG00000005885	eif4ebp2	100	94.737	ENSONIG00000018514	eif4ebp2	100	94.737	Oreochromis_niloticus
ENSMALG00000005885	eif4ebp2	98	73.684	ENSOCUG00000003475	EIF4EBP2	95	74.561	Oryctolagus_cuniculus
ENSMALG00000005885	eif4ebp2	100	92.982	ENSORLG00000022491	eif4ebp2	100	92.982	Oryzias_latipes
ENSMALG00000005885	eif4ebp2	100	92.105	ENSORLG00020005416	eif4ebp2	100	92.105	Oryzias_latipes_hni
ENSMALG00000005885	eif4ebp2	100	92.105	ENSORLG00015010580	eif4ebp2	100	92.105	Oryzias_latipes_hsok
ENSMALG00000005885	eif4ebp2	100	92.105	ENSOMEG00000005016	eif4ebp2	100	92.105	Oryzias_melastigma
ENSMALG00000005885	eif4ebp2	98	73.684	ENSOGAG00000005714	EIF4EBP2	95	74.561	Otolemur_garnettii
ENSMALG00000005885	eif4ebp2	78	72.527	ENSOARG00000006063	EIF4EBP2	95	73.626	Ovis_aries
ENSMALG00000005885	eif4ebp2	96	75.676	ENSPPAG00000022070	EIF4EBP2	93	76.577	Pan_paniscus
ENSMALG00000005885	eif4ebp2	93	65.741	ENSPPAG00000038670	-	92	66.667	Pan_paniscus
ENSMALG00000005885	eif4ebp2	96	56.881	ENSPPAG00000041572	-	94	55.357	Pan_paniscus
ENSMALG00000005885	eif4ebp2	69	67.901	ENSPPRG00000018412	EIF4EBP2	78	69.136	Panthera_pardus
ENSMALG00000005885	eif4ebp2	74	67.442	ENSPTIG00000014411	EIF4EBP2	85	68.605	Panthera_tigris_altaica
ENSMALG00000005885	eif4ebp2	98	73.684	ENSPTRG00000002590	EIF4EBP2	95	74.561	Pan_troglodytes
ENSMALG00000005885	eif4ebp2	96	56.881	ENSPTRG00000048274	-	94	55.357	Pan_troglodytes
ENSMALG00000005885	eif4ebp2	98	73.684	ENSPANG00000022261	EIF4EBP2	95	74.561	Papio_anubis
ENSMALG00000005885	eif4ebp2	96	62.832	ENSPANG00000021392	-	89	62.832	Papio_anubis
ENSMALG00000005885	eif4ebp2	96	62.162	ENSPANG00000034327	-	99	62.500	Papio_anubis
ENSMALG00000005885	eif4ebp2	100	80.702	ENSPKIG00000007681	eif4ebp2	100	80.702	Paramormyrops_kingsleyae
ENSMALG00000005885	eif4ebp2	99	76.991	ENSPSIG00000004398	EIF4EBP2	96	76.991	Pelodiscus_sinensis
ENSMALG00000005885	eif4ebp2	100	91.228	ENSPMGG00000013200	eif4ebp2	100	91.228	Periophthalmus_magnuspinnatus
ENSMALG00000005885	eif4ebp2	96	73.874	ENSPEMG00000020578	Eif4ebp2	93	74.775	Peromyscus_maniculatus_bairdii
ENSMALG00000005885	eif4ebp2	94	78.505	ENSPCIG00000015273	EIF4EBP2	85	78.505	Phascolarctos_cinereus
ENSMALG00000005885	eif4ebp2	100	95.614	ENSPFOG00000013711	eif4ebp2	100	95.614	Poecilia_formosa
ENSMALG00000005885	eif4ebp2	100	96.491	ENSPLAG00000023609	eif4ebp2	100	96.491	Poecilia_latipinna
ENSMALG00000005885	eif4ebp2	100	94.737	ENSPMEG00000021832	eif4ebp2	100	94.737	Poecilia_mexicana
ENSMALG00000005885	eif4ebp2	94	96.262	ENSPREG00000016033	eif4ebp2	96	96.262	Poecilia_reticulata
ENSMALG00000005885	eif4ebp2	98	73.684	ENSPPYG00000002371	EIF4EBP2	95	74.561	Pongo_abelii
ENSMALG00000005885	eif4ebp2	78	49.438	ENSPCAG00000014350	EIF4EBP2	92	50.495	Procavia_capensis
ENSMALG00000005885	eif4ebp2	98	73.684	ENSPCOG00000020121	EIF4EBP2	95	74.561	Propithecus_coquereli
ENSMALG00000005885	eif4ebp2	79	77.174	ENSPVAG00000000103	EIF4EBP2	93	78.261	Pteropus_vampyrus
ENSMALG00000005885	eif4ebp2	100	92.982	ENSPNYG00000011519	eif4ebp2	100	92.982	Pundamilia_nyererei
ENSMALG00000005885	eif4ebp2	100	89.474	ENSPNAG00000000572	eif4ebp2	100	89.474	Pygocentrus_nattereri
ENSMALG00000005885	eif4ebp2	96	73.874	ENSRNOG00000055391	Eif4ebp2	93	74.775	Rattus_norvegicus
ENSMALG00000005885	eif4ebp2	98	73.684	ENSRBIG00000044383	EIF4EBP2	95	74.561	Rhinopithecus_bieti
ENSMALG00000005885	eif4ebp2	98	73.684	ENSRROG00000041447	EIF4EBP2	95	74.561	Rhinopithecus_roxellana
ENSMALG00000005885	eif4ebp2	61	70.423	ENSSBOG00000021003	-	100	71.831	Saimiri_boliviensis_boliviensis
ENSMALG00000005885	eif4ebp2	100	54.386	ENSSFOG00015014062	-	85	54.386	Scleropages_formosus
ENSMALG00000005885	eif4ebp2	100	94.737	ENSSMAG00000020752	eif4ebp2	100	94.737	Scophthalmus_maximus
ENSMALG00000005885	eif4ebp2	100	98.246	ENSSDUG00000010817	eif4ebp2	100	98.246	Seriola_dumerili
ENSMALG00000005885	eif4ebp2	100	96.491	ENSSLDG00000025556	eif4ebp2	100	96.491	Seriola_lalandi_dorsalis
ENSMALG00000005885	eif4ebp2	61	70.423	ENSSARG00000000841	EIF4EBP2	100	71.831	Sorex_araneus
ENSMALG00000005885	eif4ebp2	99	76.106	ENSSPUG00000016308	EIF4EBP2	97	76.106	Sphenodon_punctatus
ENSMALG00000005885	eif4ebp2	91	98.077	ENSSPAG00000009126	eif4ebp2	88	98.077	Stegastes_partitus
ENSMALG00000005885	eif4ebp2	98	73.684	ENSSSCG00000034444	EIF4EBP2	95	74.561	Sus_scrofa
ENSMALG00000005885	eif4ebp2	61	67.143	ENSTGUG00000004591	EIF4EBP2	100	67.143	Taeniopygia_guttata
ENSMALG00000005885	eif4ebp2	97	90.090	ENSTRUG00000006895	eif4ebp2	99	90.090	Takifugu_rubripes
ENSMALG00000005885	eif4ebp2	59	69.565	ENSTBEG00000003587	EIF4EBP2	99	71.014	Tupaia_belangeri
ENSMALG00000005885	eif4ebp2	96	75.676	ENSTTRG00000003181	EIF4EBP2	93	76.577	Tursiops_truncatus
ENSMALG00000005885	eif4ebp2	98	73.684	ENSUAMG00000008214	EIF4EBP2	95	74.561	Ursus_americanus
ENSMALG00000005885	eif4ebp2	61	70.833	ENSUMAG00000007968	EIF4EBP2	64	72.222	Ursus_maritimus
ENSMALG00000005885	eif4ebp2	59	55.224	ENSVPAG00000002634	-	97	63.768	Vicugna_pacos
ENSMALG00000005885	eif4ebp2	98	73.684	ENSVVUG00000021616	EIF4EBP2	95	74.561	Vulpes_vulpes
ENSMALG00000005885	eif4ebp2	100	73.684	ENSXETG00000025651	eif4ebp2	97	73.684	Xenopus_tropicalis
ENSMALG00000005885	eif4ebp2	100	94.737	ENSXCOG00000013288	eif4ebp2	100	94.737	Xiphophorus_couchianus
ENSMALG00000005885	eif4ebp2	100	95.614	ENSXMAG00000003157	eif4ebp2	100	95.614	Xiphophorus_maculatus