EuRBPDB

Description

RBDs

Transcripts

Gene Model

PPI

Paralogs

Orthologs

Description

Ensembl ID: ENSOGAG00000001415 (Gene tree)
Gene ID: -
Gene Symbol: LARP4
Alias: -
Full Name: La ribonucleoprotein domain family member 4
Gene Type: protein_coding
Species: Otolemur_garnettii
Status: putative
Strand: Plus strand
Length: 40423 bases
Position: chrGL873562.1:4809689-4850111
Accession: 24320
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
ENSOGAP00000001260	La	PF05383.17	3.4e-17	1	1

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
ENSOGAT00000001416	LARP4-201	1554	-	ENSOGAP00000001260	518 (aa)	-	H0WIJ5

Gene Model

Click here to download ENSOGAG00000001415's gene model file

Protein-Protein Interaction (PPI)

Clik here to download ENSOGAG00000001415's network

Paralogs

Ensembl ID	Gene Symbol	Coverage	Identiy	Paralog	Gene Symbol	Coverage	Identiy
ENSOGAG00000001415	LARP4	99	39.817	ENSOGAG00000011540	LARP4B	75	39.083

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
ENSOGAG00000001415	LARP4	100	89.403	ENSG00000161813	LARP4	100	96.800	Homo_sapiens
ENSOGAG00000001415	LARP4	98	52.008	ENSAPOG00000021331	larp4ab	68	50.476	Acanthochromis_polyacanthus
ENSOGAG00000001415	LARP4	100	87.838	ENSAMEG00000001275	LARP4	71	87.838	Ailuropoda_melanoleuca
ENSOGAG00000001415	LARP4	99	50.847	ENSACIG00000001198	larp4aa	93	72.619	Amphilophus_citrinellus
ENSOGAG00000001415	LARP4	98	51.619	ENSAOCG00000006387	larp4ab	68	50.000	Amphiprion_ocellaris
ENSOGAG00000001415	LARP4	99	54.699	ENSAOCG00000002408	larp4aa	53	63.265	Amphiprion_ocellaris
ENSOGAG00000001415	LARP4	99	54.887	ENSAPEG00000019058	larp4aa	53	63.265	Amphiprion_percula
ENSOGAG00000001415	LARP4	72	55.080	ENSAPEG00000015441	larp4ab	98	55.080	Amphiprion_percula
ENSOGAG00000001415	LARP4	98	47.834	ENSATEG00000001018	larp4ab	68	47.932	Anabas_testudineus
ENSOGAG00000001415	LARP4	100	72.433	ENSAPLG00000013661	LARP4	71	71.863	Anas_platyrhynchos
ENSOGAG00000001415	LARP4	99	69.714	ENSACAG00000017133	LARP4	69	69.143	Anolis_carolinensis
ENSOGAG00000001415	LARP4	100	88.632	ENSANAG00000029342	LARP4	71	88.632	Aotus_nancymaae
ENSOGAG00000001415	LARP4	72	53.766	ENSACLG00000000777	larp4ab	98	53.766	Astatotilapia_calliptera
ENSOGAG00000001415	LARP4	99	49.905	ENSAMXG00000001515	larp4ab	66	49.442	Astyanax_mexicanus
ENSOGAG00000001415	LARP4	99	55.556	ENSAMXG00000036493	larp4aa	63	55.179	Astyanax_mexicanus
ENSOGAG00000001415	LARP4	100	86.364	ENSBTAG00000017164	LARP4	71	86.364	Bos_taurus
ENSOGAG00000001415	LARP4	100	88.031	ENSCJAG00000021131	LARP4	71	88.031	Callithrix_jacchus
ENSOGAG00000001415	LARP4	100	88.610	ENSCAFG00000008174	LARP4	71	88.610	Canis_familiaris
ENSOGAG00000001415	LARP4	100	88.803	ENSCAFG00020014554	LARP4	73	88.713	Canis_lupus_dingo
ENSOGAG00000001415	LARP4	100	86.553	ENSCHIG00000025965	LARP4	71	86.553	Capra_hircus
ENSOGAG00000001415	LARP4	100	89.189	ENSTSYG00000004342	LARP4	71	89.189	Carlito_syrichta
ENSOGAG00000001415	LARP4	99	63.301	ENSCAPG00000012942	LARP4	68	65.820	Cavia_aperea
ENSOGAG00000001415	LARP4	99	78.101	ENSCPOG00000020519	LARP4	70	78.101	Cavia_porcellus
ENSOGAG00000001415	LARP4	100	87.452	ENSCCAG00000024439	LARP4	71	87.452	Cebus_capucinus
ENSOGAG00000001415	LARP4	100	87.452	ENSCATG00000044528	LARP4	71	87.452	Cercocebus_atys
ENSOGAG00000001415	LARP4	99	85.603	ENSCLAG00000001829	LARP4	70	85.603	Chinchilla_lanigera
ENSOGAG00000001415	LARP4	100	73.574	ENSCPBG00000019407	LARP4	64	73.004	Chrysemys_picta_bellii
ENSOGAG00000001415	LARP4	100	78.958	ENSCANG00000033097	-	81	77.451	Colobus_angolensis_palliatus
ENSOGAG00000001415	LARP4	100	89.768	ENSCANG00000029159	-	71	89.768	Colobus_angolensis_palliatus
ENSOGAG00000001415	LARP4	99	84.825	ENSCGRG00001010818	Larp4	70	84.825	Cricetulus_griseus_chok1gshd
ENSOGAG00000001415	LARP4	99	84.825	ENSCGRG00000009382	Larp4	71	84.795	Cricetulus_griseus_crigri
ENSOGAG00000001415	LARP4	99	54.080	ENSCSEG00000001963	larp4aa	51	62.500	Cynoglossus_semilaevis
ENSOGAG00000001415	LARP4	99	50.471	ENSDARG00000074979	larp4ab	91	73.373	Danio_rerio
ENSOGAG00000001415	LARP4	99	52.757	ENSDARG00000078401	larp4aa	65	51.661	Danio_rerio
ENSOGAG00000001415	LARP4	100	88.610	ENSDNOG00000038525	LARP4	71	88.610	Dasypus_novemcinctus
ENSOGAG00000001415	LARP4	99	86.434	ENSDORG00000004013	Larp4	70	86.434	Dipodomys_ordii
ENSOGAG00000001415	LARP4	98	41.103	ENSEBUG00000004426	LARP4	92	40.747	Eptatretus_burgeri
ENSOGAG00000001415	LARP4	89	40.779	ENSEBUG00000010394	LARP4	68	43.326	Eptatretus_burgeri
ENSOGAG00000001415	LARP4	100	89.382	ENSEASG00005004356	LARP4	98	89.867	Equus_asinus_asinus
ENSOGAG00000001415	LARP4	100	89.768	ENSECAG00000012479	LARP4	71	89.768	Equus_caballus
ENSOGAG00000001415	LARP4	99	48.204	ENSELUG00000000395	larp4aa	56	63.265	Esox_lucius
ENSOGAG00000001415	LARP4	98	52.273	ENSELUG00000018650	larp4ab	51	68.511	Esox_lucius
ENSOGAG00000001415	LARP4	100	88.803	ENSFCAG00000031897	LARP4	71	88.803	Felis_catus
ENSOGAG00000001415	LARP4	100	71.893	ENSFALG00000002114	LARP4	79	71.319	Ficedula_albicollis
ENSOGAG00000001415	LARP4	99	85.659	ENSFDAG00000006406	-	71	85.659	Fukomys_damarensis
ENSOGAG00000001415	LARP4	98	49.341	ENSFHEG00000013854	larp4ab	68	48.956	Fundulus_heteroclitus
ENSOGAG00000001415	LARP4	99	52.166	ENSGMOG00000008246	larp4aa	66	50.943	Gadus_morhua
ENSOGAG00000001415	LARP4	65	47.550	ENSGMOG00000009536	-	50	47.262	Gadus_morhua
ENSOGAG00000001415	LARP4	99	72.414	ENSGALG00000042122	LARP4	69	71.839	Gallus_gallus
ENSOGAG00000001415	LARP4	52	70.849	ENSGAFG00000004280	larp4aa	74	70.849	Gambusia_affinis
ENSOGAG00000001415	LARP4	98	49.159	ENSGAFG00000021777	larp4ab	69	48.411	Gambusia_affinis
ENSOGAG00000001415	LARP4	98	50.000	ENSGACG00000000680	larp4ab	75	47.505	Gasterosteus_aculeatus
ENSOGAG00000001415	LARP4	99	54.302	ENSGACG00000008381	larp4aa	98	60.158	Gasterosteus_aculeatus
ENSOGAG00000001415	LARP4	99	72.797	ENSGAGG00000006969	LARP4	71	72.222	Gopherus_agassizii
ENSOGAG00000001415	LARP4	100	89.189	ENSGGOG00000015230	LARP4	99	90.027	Gorilla_gorilla
ENSOGAG00000001415	LARP4	73	57.881	ENSHBUG00000023074	larp4aa	50	57.881	Haplochromis_burtoni
ENSOGAG00000001415	LARP4	99	87.160	ENSHGLG00000005513	-	78	87.160	Heterocephalus_glaber_female
ENSOGAG00000001415	LARP4	87	69.469	ENSHGLG00000010979	-	92	71.136	Heterocephalus_glaber_female
ENSOGAG00000001415	LARP4	82	84.038	ENSHGLG00000013875	-	71	82.394	Heterocephalus_glaber_female
ENSOGAG00000001415	LARP4	99	82.171	ENSHGLG00100011678	-	72	80.814	Heterocephalus_glaber_male
ENSOGAG00000001415	LARP4	66	75.731	ENSHGLG00100000622	-	97	75.731	Heterocephalus_glaber_male
ENSOGAG00000001415	LARP4	87	69.469	ENSHGLG00100002832	-	92	71.136	Heterocephalus_glaber_male
ENSOGAG00000001415	LARP4	99	86.486	ENSHGLG00100016984	-	72	86.486	Heterocephalus_glaber_male
ENSOGAG00000001415	LARP4	98	45.437	ENSHCOG00000020857	larp4ab	69	45.437	Hippocampus_comes
ENSOGAG00000001415	LARP4	99	52.642	ENSHCOG00000015951	larp4aa	65	53.484	Hippocampus_comes
ENSOGAG00000001415	LARP4	99	52.612	ENSIPUG00000007514	larp4aa	64	51.493	Ictalurus_punctatus
ENSOGAG00000001415	LARP4	100	50.766	ENSIPUG00000021659	larp4ab	74	50.288	Ictalurus_punctatus
ENSOGAG00000001415	LARP4	99	88.953	ENSSTOG00000003553	LARP4	71	88.953	Ictidomys_tridecemlineatus
ENSOGAG00000001415	LARP4	71	64.439	ENSJJAG00000014649	-	95	85.581	Jaculus_jaculus
ENSOGAG00000001415	LARP4	98	50.473	ENSKMAG00000018556	larp4ab	70	49.716	Kryptolebias_marmoratus
ENSOGAG00000001415	LARP4	99	53.846	ENSKMAG00000006800	larp4aa	65	49.468	Kryptolebias_marmoratus
ENSOGAG00000001415	LARP4	98	49.146	ENSLBEG00000019306	larp4ab	69	49.146	Labrus_bergylta
ENSOGAG00000001415	LARP4	99	60.935	ENSLACG00000019090	LARP4	71	59.626	Latimeria_chalumnae
ENSOGAG00000001415	LARP4	99	56.604	ENSLOCG00000007113	larp4ab	66	55.198	Lepisosteus_oculatus
ENSOGAG00000001415	LARP4	100	88.417	ENSLAFG00000005160	LARP4	71	88.417	Loxodonta_africana
ENSOGAG00000001415	LARP4	100	90.154	ENSMFAG00000043681	LARP4	73	90.154	Macaca_fascicularis
ENSOGAG00000001415	LARP4	100	90.751	ENSMMUG00000008172	LARP4	71	90.751	Macaca_mulatta
ENSOGAG00000001415	LARP4	90	90.792	ENSMNEG00000034602	LARP4	71	91.091	Macaca_nemestrina
ENSOGAG00000001415	LARP4	81	70.089	ENSMNEG00000037799	-	99	76.216	Macaca_nemestrina
ENSOGAG00000001415	LARP4	100	90.347	ENSMLEG00000039661	LARP4	71	90.347	Mandrillus_leucophaeus
ENSOGAG00000001415	LARP4	98	49.813	ENSMAMG00000002616	larp4ab	69	49.813	Mastacembelus_armatus
ENSOGAG00000001415	LARP4	99	53.296	ENSMAMG00000011206	larp4aa	64	53.759	Mastacembelus_armatus
ENSOGAG00000001415	LARP4	72	53.766	ENSMZEG00005007771	larp4ab	98	53.766	Maylandia_zebra
ENSOGAG00000001415	LARP4	99	72.605	ENSMGAG00000010598	LARP4	68	72.031	Meleagris_gallopavo
ENSOGAG00000001415	LARP4	99	84.795	ENSMICG00000030156	-	87	86.772	Microcebus_murinus
ENSOGAG00000001415	LARP4	82	89.882	ENSMICG00000012388	-	66	89.882	Microcebus_murinus
ENSOGAG00000001415	LARP4	99	84.302	ENSMOCG00000020635	Larp4	71	84.302	Microtus_ochrogaster
ENSOGAG00000001415	LARP4	98	48.077	ENSMMOG00000021861	larp4ab	91	53.533	Mola_mola
ENSOGAG00000001415	LARP4	99	51.673	ENSMMOG00000004785	larp4aa	66	50.743	Mola_mola
ENSOGAG00000001415	LARP4	52	67.897	ENSMALG00000018631	larp4aa	98	67.708	Monopterus_albus
ENSOGAG00000001415	LARP4	99	75.486	MGP_CAROLIEiJ_G0020281	Larp4	84	83.283	Mus_caroli
ENSOGAG00000001415	LARP4	99	84.241	ENSMUSG00000023025	Larp4	83	84.241	Mus_musculus
ENSOGAG00000001415	LARP4	100	84.363	MGP_PahariEiJ_G0020284	Larp4	71	84.363	Mus_pahari
ENSOGAG00000001415	LARP4	100	87.645	ENSMPUG00000008706	LARP4	69	87.645	Mustela_putorius_furo
ENSOGAG00000001415	LARP4	100	86.680	ENSMLUG00000012513	-	70	86.680	Myotis_lucifugus
ENSOGAG00000001415	LARP4	99	83.721	ENSNGAG00000001577	-	71	83.721	Nannospalax_galili
ENSOGAG00000001415	LARP4	72	52.727	ENSNBRG00000015675	larp4ab	94	61.111	Neolamprologus_brichardi
ENSOGAG00000001415	LARP4	100	89.575	ENSNLEG00000017798	LARP4	71	89.575	Nomascus_leucogenys
ENSOGAG00000001415	LARP4	76	85.315	ENSOPRG00000004592	-	64	85.315	Ochotona_princeps
ENSOGAG00000001415	LARP4	99	86.160	ENSODEG00000009770	LARP4	72	86.160	Octodon_degus
ENSOGAG00000001415	LARP4	98	48.099	ENSONIG00000019124	larp4ab	70	47.438	Oreochromis_niloticus
ENSOGAG00000001415	LARP4	99	53.757	ENSONIG00000011895	larp4aa	67	54.913	Oreochromis_niloticus
ENSOGAG00000001415	LARP4	99	70.690	ENSOANG00000004373	LARP4	93	70.690	Ornithorhynchus_anatinus
ENSOGAG00000001415	LARP4	99	86.873	ENSOCUG00000026872	-	71	84.363	Oryctolagus_cuniculus
ENSOGAG00000001415	LARP4	99	85.825	ENSOCUG00000021417	-	70	85.825	Oryctolagus_cuniculus
ENSOGAG00000001415	LARP4	99	54.876	ENSORLG00000006758	larp4aa	64	53.802	Oryzias_latipes
ENSOGAG00000001415	LARP4	99	55.067	ENSORLG00020010258	larp4aa	64	53.257	Oryzias_latipes_hni
ENSOGAG00000001415	LARP4	99	54.857	ENSORLG00015002292	larp4aa	64	53.788	Oryzias_latipes_hsok
ENSOGAG00000001415	LARP4	98	55.556	ENSOMEG00000019749	larp4aa	96	62.295	Oryzias_melastigma
ENSOGAG00000001415	LARP4	100	86.553	ENSOARG00000017702	LARP4	71	86.553	Ovis_aries
ENSOGAG00000001415	LARP4	87	70.133	ENSPPAG00000039122	-	94	74.390	Pan_paniscus
ENSOGAG00000001415	LARP4	85	88.182	ENSPPAG00000035100	LARP4	65	88.182	Pan_paniscus
ENSOGAG00000001415	LARP4	100	88.996	ENSPPRG00000004522	LARP4	71	88.996	Panthera_pardus
ENSOGAG00000001415	LARP4	100	88.996	ENSPTIG00000015617	LARP4	71	88.996	Panthera_tigris_altaica
ENSOGAG00000001415	LARP4	87	69.690	ENSPTRG00000045907	-	97	75.543	Pan_troglodytes
ENSOGAG00000001415	LARP4	100	89.189	ENSPTRG00000004935	LARP4	71	89.189	Pan_troglodytes
ENSOGAG00000001415	LARP4	100	90.944	ENSPANG00000022964	LARP4	71	90.944	Papio_anubis
ENSOGAG00000001415	LARP4	99	54.545	ENSPKIG00000005950	larp4ab	68	51.381	Paramormyrops_kingsleyae
ENSOGAG00000001415	LARP4	99	70.441	ENSPSIG00000015867	LARP4	70	70.441	Pelodiscus_sinensis
ENSOGAG00000001415	LARP4	70	50.938	ENSPMGG00000002719	larp4ab	96	51.475	Periophthalmus_magnuspinnatus
ENSOGAG00000001415	LARP4	99	76.346	ENSPEMG00000018433	Larp4	62	85.468	Peromyscus_maniculatus_bairdii
ENSOGAG00000001415	LARP4	100	76.616	ENSPCIG00000000384	LARP4	71	76.046	Phascolarctos_cinereus
ENSOGAG00000001415	LARP4	99	52.919	ENSPFOG00000013148	larp4aa	74	53.392	Poecilia_formosa
ENSOGAG00000001415	LARP4	98	49.809	ENSPFOG00000009764	larp4ab	69	49.430	Poecilia_formosa
ENSOGAG00000001415	LARP4	99	52.919	ENSPLAG00000013626	larp4aa	64	50.814	Poecilia_latipinna
ENSOGAG00000001415	LARP4	98	49.618	ENSPLAG00000008502	larp4ab	69	49.240	Poecilia_latipinna
ENSOGAG00000001415	LARP4	99	52.820	ENSPMEG00000014271	larp4aa	64	50.814	Poecilia_mexicana
ENSOGAG00000001415	LARP4	98	50.093	ENSPMEG00000002720	larp4ab	69	49.533	Poecilia_mexicana
ENSOGAG00000001415	LARP4	72	55.643	ENSPREG00000016263	larp4ab	99	56.863	Poecilia_reticulata
ENSOGAG00000001415	LARP4	99	54.766	ENSPREG00000022597	larp4aa	65	50.633	Poecilia_reticulata
ENSOGAG00000001415	LARP4	99	89.729	ENSPPYG00000004512	-	99	89.729	Pongo_abelii
ENSOGAG00000001415	LARP4	90	78.632	ENSPCAG00000003671	LARP4	69	78.632	Procavia_capensis
ENSOGAG00000001415	LARP4	82	98.182	ENSPCOG00000026291	LARP4	66	98.182	Propithecus_coquereli
ENSOGAG00000001415	LARP4	90	85.653	ENSPVAG00000016840	LARP4	66	85.653	Pteropus_vampyrus
ENSOGAG00000001415	LARP4	99	52.075	ENSPNAG00000014478	larp4ab	70	51.017	Pygocentrus_nattereri
ENSOGAG00000001415	LARP4	99	54.631	ENSPNAG00000024770	larp4aa	67	54.682	Pygocentrus_nattereri
ENSOGAG00000001415	LARP4	99	90.541	ENSRBIG00000038857	-	98	89.773	Rhinopithecus_bieti
ENSOGAG00000001415	LARP4	83	82.286	ENSRBIG00000036890	-	62	82.286	Rhinopithecus_bieti
ENSOGAG00000001415	LARP4	100	90.751	ENSRROG00000042764	-	98	89.773	Rhinopithecus_roxellana
ENSOGAG00000001415	LARP4	100	76.834	ENSRROG00000025487	-	89	74.332	Rhinopithecus_roxellana
ENSOGAG00000001415	LARP4	100	87.645	ENSSBOG00000026650	LARP4	71	87.645	Saimiri_boliviensis_boliviensis
ENSOGAG00000001415	LARP4	65	79.883	ENSSHAG00000013246	-	99	79.941	Sarcophilus_harrisii
ENSOGAG00000001415	LARP4	99	52.008	ENSSFOG00015014096	larp4ab	68	52.390	Scleropages_formosus
ENSOGAG00000001415	LARP4	99	56.538	ENSSFOG00015003868	larp4aa	69	54.615	Scleropages_formosus
ENSOGAG00000001415	LARP4	98	52.354	ENSSMAG00000002838	larp4ab	69	52.166	Scophthalmus_maximus
ENSOGAG00000001415	LARP4	98	51.128	ENSSDUG00000006655	larp4ab	69	51.316	Seriola_dumerili
ENSOGAG00000001415	LARP4	98	50.564	ENSSLDG00000001791	larp4ab	69	50.657	Seriola_lalandi_dorsalis
ENSOGAG00000001415	LARP4	100	86.127	ENSSARG00000012469	LARP4	72	86.127	Sorex_araneus
ENSOGAG00000001415	LARP4	99	71.456	ENSSPUG00000016552	LARP4	69	71.456	Sphenodon_punctatus
ENSOGAG00000001415	LARP4	98	52.099	ENSSPAG00000002172	larp4ab	69	50.575	Stegastes_partitus
ENSOGAG00000001415	LARP4	99	53.107	ENSSPAG00000007222	larp4aa	98	59.249	Stegastes_partitus
ENSOGAG00000001415	LARP4	100	88.439	ENSSSCG00000028516	LARP4	71	88.439	Sus_scrofa
ENSOGAG00000001415	LARP4	99	71.236	ENSTGUG00000003479	LARP4	70	71.236	Taeniopygia_guttata
ENSOGAG00000001415	LARP4	98	48.674	ENSTRUG00000007206	larp4ab	70	48.485	Takifugu_rubripes
ENSOGAG00000001415	LARP4	70	56.474	ENSTRUG00000015939	larp4aa	100	56.474	Takifugu_rubripes
ENSOGAG00000001415	LARP4	99	51.985	ENSTNIG00000015109	larp4aa	69	51.698	Tetraodon_nigroviridis
ENSOGAG00000001415	LARP4	99	48.315	ENSTNIG00000014571	larp4ab	73	48.023	Tetraodon_nigroviridis
ENSOGAG00000001415	LARP4	100	73.359	ENSTBEG00000005167	LARP4	72	73.359	Tupaia_belangeri
ENSOGAG00000001415	LARP4	99	86.408	ENSTTRG00000003764	LARP4	70	86.408	Tursiops_truncatus
ENSOGAG00000001415	LARP4	100	88.610	ENSUAMG00000013764	LARP4	98	88.864	Ursus_americanus
ENSOGAG00000001415	LARP4	100	88.417	ENSUMAG00000008473	LARP4	71	88.417	Ursus_maritimus
ENSOGAG00000001415	LARP4	100	87.048	ENSVPAG00000010136	LARP4	71	87.048	Vicugna_pacos
ENSOGAG00000001415	LARP4	100	88.996	ENSVVUG00000024717	LARP4	72	88.996	Vulpes_vulpes
ENSOGAG00000001415	LARP4	90	62.185	ENSXETG00000007283	larp4	70	62.185	Xenopus_tropicalis
ENSOGAG00000001415	LARP4	72	54.856	ENSXCOG00000019635	larp4ab	99	55.344	Xiphophorus_couchianus
ENSOGAG00000001415	LARP4	99	50.755	ENSXCOG00000006387	larp4aa	63	50.000	Xiphophorus_couchianus
ENSOGAG00000001415	LARP4	98	50.857	ENSXMAG00000002156	larp4ab	69	50.095	Xiphophorus_maculatus
ENSOGAG00000001415	LARP4	99	54.275	ENSXMAG00000010519	larp4aa	65	52.952	Xiphophorus_maculatus