EuRBPDB

Description

RBDs

Transcripts

Gene Model

PPI

Paralogs

Orthologs

Description

Ensembl ID: ENSPREG00000021082 (Gene tree)
Gene ID: 103481195
Gene Symbol: CRACR2A (1 of many)
Alias: -
Full Name: ras and EF-hand domain-containing protein homolog
Gene Type: protein_coding
Species: Poecilia_reticulata
Status: putative
Strand: Plus strand
Length: 11118 bases
Position: chrLG19:3951281-3962398
Accession: 103481195
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
ENSPREP00000031121	MMR_HSR1	PF01926.23	2.5e-08	1	1

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
ENSPRET00000031474	CRACR2A (1 of many)-201	2327	XM_008436518	ENSPREP00000031121	717 (aa)	XP_008434740	UPI0004A45195

Gene Model

Click here to download ENSPREG00000021082's gene model file

Protein-Protein Interaction (PPI)

Clik here to download ENSPREG00000021082's network

Paralogs

Ensembl ID	Gene Symbol	Coverage	Identiy	Paralog	Gene Symbol	Coverage	Identiy
ENSPREG00000021082	CRACR2A	72	39.487	ENSPREG00000018728	rasef	61	39.487
ENSPREG00000021082	CRACR2A	96	54.762	ENSPREG00000002593	cracr2ab	98	55.556

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
ENSPREG00000021082	CRACR2A	96	52.528	ENSG00000130038	CRACR2A	90	59.218	Homo_sapiens
ENSPREG00000021082	CRACR2A	76	50.777	ENSAPOG00000009642	cracr2b	85	47.926	Acanthochromis_polyacanthus
ENSPREG00000021082	CRACR2A	90	54.138	ENSAPOG00000013779	cracr2ab	99	54.138	Acanthochromis_polyacanthus
ENSPREG00000021082	CRACR2A	70	55.039	ENSAMEG00000005041	-	97	69.880	Ailuropoda_melanoleuca
ENSPREG00000021082	CRACR2A	95	55.143	ENSACIG00000004009	cracr2ab	97	55.920	Amphilophus_citrinellus
ENSPREG00000021082	CRACR2A	100	71.034	ENSACIG00000010099	CRACR2A	100	71.034	Amphilophus_citrinellus
ENSPREG00000021082	CRACR2A	93	35.863	ENSAOCG00000019493	cracr2b	89	36.269	Amphiprion_ocellaris
ENSPREG00000021082	CRACR2A	100	73.806	ENSAOCG00000008147	-	100	73.806	Amphiprion_ocellaris
ENSPREG00000021082	CRACR2A	95	54.675	ENSAOCG00000023189	cracr2ab	98	55.065	Amphiprion_ocellaris
ENSPREG00000021082	CRACR2A	93	36.310	ENSAPEG00000021843	cracr2b	90	36.567	Amphiprion_percula
ENSPREG00000021082	CRACR2A	95	54.339	ENSAPEG00000006383	cracr2ab	99	54.688	Amphiprion_percula
ENSPREG00000021082	CRACR2A	100	73.806	ENSAPEG00000002244	CRACR2A	100	73.806	Amphiprion_percula
ENSPREG00000021082	CRACR2A	100	72.230	ENSATEG00000017411	CRACR2A	100	72.230	Anabas_testudineus
ENSPREG00000021082	CRACR2A	78	48.624	ENSATEG00000010344	cracr2b	86	48.624	Anabas_testudineus
ENSPREG00000021082	CRACR2A	87	66.477	ENSATEG00000018003	cracr2ab	82	66.477	Anabas_testudineus
ENSPREG00000021082	CRACR2A	94	38.659	ENSACAG00000015413	CRACR2B	94	38.659	Anolis_carolinensis
ENSPREG00000021082	CRACR2A	96	53.835	ENSANAG00000036979	CRACR2A	95	53.835	Aotus_nancymaae
ENSPREG00000021082	CRACR2A	95	53.604	ENSACLG00000023670	cracr2ab	98	53.735	Astatotilapia_calliptera
ENSPREG00000021082	CRACR2A	100	71.664	ENSACLG00000007547	CRACR2A	100	71.664	Astatotilapia_calliptera
ENSPREG00000021082	CRACR2A	94	34.911	ENSACLG00000013916	cracr2b	89	34.911	Astatotilapia_calliptera
ENSPREG00000021082	CRACR2A	57	48.081	ENSAMXG00000010368	cracr2ab	99	48.423	Astyanax_mexicanus
ENSPREG00000021082	CRACR2A	94	53.094	ENSBTAG00000018940	CRACR2A	94	53.094	Bos_taurus
ENSPREG00000021082	CRACR2A	96	53.371	ENSCJAG00000018917	CRACR2A	95	53.814	Callithrix_jacchus
ENSPREG00000021082	CRACR2A	94	53.372	ENSCAFG00000015404	CRACR2A	94	53.802	Canis_familiaris
ENSPREG00000021082	CRACR2A	94	53.372	ENSCAFG00020014835	CRACR2A	94	53.802	Canis_lupus_dingo
ENSPREG00000021082	CRACR2A	94	54.388	ENSCHIG00000018141	CRACR2A	94	54.676	Capra_hircus
ENSPREG00000021082	CRACR2A	94	52.511	ENSTSYG00000014194	CRACR2A	94	52.511	Carlito_syrichta
ENSPREG00000021082	CRACR2A	86	46.009	ENSCAPG00000006188	CRACR2A	100	46.009	Cavia_aperea
ENSPREG00000021082	CRACR2A	94	50.925	ENSCPOG00000020119	CRACR2A	95	50.925	Cavia_porcellus
ENSPREG00000021082	CRACR2A	96	54.278	ENSCCAG00000019735	CRACR2A	95	54.558	Cebus_capucinus
ENSPREG00000021082	CRACR2A	96	52.859	ENSCATG00000040964	CRACR2A	95	53.738	Cercocebus_atys
ENSPREG00000021082	CRACR2A	94	51.433	ENSCLAG00000013246	CRACR2A	95	50.989	Chinchilla_lanigera
ENSPREG00000021082	CRACR2A	96	53.147	ENSCSAG00000011199	CRACR2A	95	53.727	Chlorocebus_sabaeus
ENSPREG00000021082	CRACR2A	95	42.614	ENSCPBG00000007146	CRACR2B	91	42.756	Chrysemys_picta_bellii
ENSPREG00000021082	CRACR2A	93	55.637	ENSCPBG00000009731	CRACR2A	93	55.637	Chrysemys_picta_bellii
ENSPREG00000021082	CRACR2A	96	53.296	ENSCANG00000032776	CRACR2A	95	53.717	Colobus_angolensis_palliatus
ENSPREG00000021082	CRACR2A	94	52.086	ENSCGRG00001008752	-	94	52.086	Cricetulus_griseus_chok1gshd
ENSPREG00000021082	CRACR2A	94	51.511	ENSCGRG00000015501	-	94	51.511	Cricetulus_griseus_crigri
ENSPREG00000021082	CRACR2A	84	67.232	ENSCSEG00000008307	cracr2ab	86	63.566	Cynoglossus_semilaevis
ENSPREG00000021082	CRACR2A	100	64.404	ENSCSEG00000003612	CRACR2A	100	64.404	Cynoglossus_semilaevis
ENSPREG00000021082	CRACR2A	72	49.468	ENSCSEG00000007618	cracr2b	80	47.000	Cynoglossus_semilaevis
ENSPREG00000021082	CRACR2A	96	55.334	ENSCVAG00000009954	cracr2ab	98	55.334	Cyprinodon_variegatus
ENSPREG00000021082	CRACR2A	95	81.449	ENSCVAG00000013782	CRACR2A	99	81.449	Cyprinodon_variegatus
ENSPREG00000021082	CRACR2A	94	53.774	ENSDARG00000045758	cracr2ab	95	53.774	Danio_rerio
ENSPREG00000021082	CRACR2A	100	61.937	ENSDARG00000068231	CRACR2A	100	61.801	Danio_rerio
ENSPREG00000021082	CRACR2A	97	50.205	ENSDNOG00000043722	CRACR2A	97	50.752	Dasypus_novemcinctus
ENSPREG00000021082	CRACR2A	94	52.035	ENSEASG00005017095	CRACR2A	94	52.035	Equus_asinus_asinus
ENSPREG00000021082	CRACR2A	94	53.458	ENSECAG00000034127	CRACR2A	94	53.458	Equus_caballus
ENSPREG00000021082	CRACR2A	91	63.708	ENSELUG00000001781	cracr2ab	77	63.708	Esox_lucius
ENSPREG00000021082	CRACR2A	100	62.687	ENSELUG00000018918	CRACR2A	100	62.687	Esox_lucius
ENSPREG00000021082	CRACR2A	94	53.314	ENSFCAG00000015741	CRACR2A	94	53.602	Felis_catus
ENSPREG00000021082	CRACR2A	95	43.706	ENSFALG00000010481	CRACR2B	95	43.706	Ficedula_albicollis
ENSPREG00000021082	CRACR2A	97	52.448	ENSFALG00000004782	CRACR2A	99	52.448	Ficedula_albicollis
ENSPREG00000021082	CRACR2A	98	49.533	ENSFDAG00000013877	CRACR2A	98	49.666	Fukomys_damarensis
ENSPREG00000021082	CRACR2A	95	47.706	ENSFHEG00000021525	cracr2ab	98	53.403	Fundulus_heteroclitus
ENSPREG00000021082	CRACR2A	100	93.750	ENSGAFG00000014607	CRACR2A	100	93.750	Gambusia_affinis
ENSPREG00000021082	CRACR2A	75	49.474	ENSGAFG00000020778	cracr2b	79	49.474	Gambusia_affinis
ENSPREG00000021082	CRACR2A	95	51.554	ENSGAFG00000008771	cracr2ab	98	51.554	Gambusia_affinis
ENSPREG00000021082	CRACR2A	93	56.955	ENSGAGG00000002264	CRACR2A	93	56.955	Gopherus_agassizii
ENSPREG00000021082	CRACR2A	96	52.528	ENSGGOG00000010289	CRACR2A	95	53.107	Gorilla_gorilla
ENSPREG00000021082	CRACR2A	95	53.604	ENSHBUG00000018719	cracr2ab	98	53.735	Haplochromis_burtoni
ENSPREG00000021082	CRACR2A	100	71.527	ENSHBUG00000017137	CRACR2A	100	71.527	Haplochromis_burtoni
ENSPREG00000021082	CRACR2A	94	52.865	ENSHGLG00000003776	CRACR2A	95	52.865	Heterocephalus_glaber_female
ENSPREG00000021082	CRACR2A	94	52.865	ENSHGLG00100005482	CRACR2A	95	52.865	Heterocephalus_glaber_male
ENSPREG00000021082	CRACR2A	96	57.759	ENSHCOG00000004942	cracr2ab	99	57.759	Hippocampus_comes
ENSPREG00000021082	CRACR2A	99	66.713	ENSHCOG00000006842	CRACR2A	99	66.713	Hippocampus_comes
ENSPREG00000021082	CRACR2A	78	44.541	ENSIPUG00000013598	cracr2b	85	44.541	Ictalurus_punctatus
ENSPREG00000021082	CRACR2A	97	53.425	ENSIPUG00000017914	cracr2ab	97	53.425	Ictalurus_punctatus
ENSPREG00000021082	CRACR2A	100	61.976	ENSIPUG00000015191	CRACR2A	100	61.976	Ictalurus_punctatus
ENSPREG00000021082	CRACR2A	96	51.836	ENSSTOG00000021197	CRACR2A	95	51.836	Ictidomys_tridecemlineatus
ENSPREG00000021082	CRACR2A	94	49.000	ENSJJAG00000014186	-	93	49.000	Jaculus_jaculus
ENSPREG00000021082	CRACR2A	95	53.448	ENSKMAG00000016358	cracr2ab	97	53.448	Kryptolebias_marmoratus
ENSPREG00000021082	CRACR2A	99	74.028	ENSKMAG00000021906	CRACR2A	100	73.098	Kryptolebias_marmoratus
ENSPREG00000021082	CRACR2A	95	55.702	ENSLBEG00000018669	cracr2ab	97	55.848	Labrus_bergylta
ENSPREG00000021082	CRACR2A	94	72.117	ENSLBEG00000010713	CRACR2A	100	70.014	Labrus_bergylta
ENSPREG00000021082	CRACR2A	93	32.650	ENSLBEG00000014261	cracr2b	91	32.650	Labrus_bergylta
ENSPREG00000021082	CRACR2A	94	39.045	ENSLACG00000007778	-	99	39.045	Latimeria_chalumnae
ENSPREG00000021082	CRACR2A	94	40.145	ENSLACG00000009026	-	99	40.145	Latimeria_chalumnae
ENSPREG00000021082	CRACR2A	100	63.073	ENSLOCG00000016848	cracr2ab	99	63.798	Lepisosteus_oculatus
ENSPREG00000021082	CRACR2A	94	53.770	ENSMFAG00000036067	CRACR2A	95	54.020	Macaca_fascicularis
ENSPREG00000021082	CRACR2A	96	52.859	ENSMMUG00000023583	CRACR2A	95	53.597	Macaca_mulatta
ENSPREG00000021082	CRACR2A	96	52.999	ENSMNEG00000031773	CRACR2A	95	53.879	Macaca_nemestrina
ENSPREG00000021082	CRACR2A	96	52.720	ENSMLEG00000044159	CRACR2A	95	53.597	Mandrillus_leucophaeus
ENSPREG00000021082	CRACR2A	99	72.361	ENSMAMG00000001270	CRACR2A	100	71.760	Mastacembelus_armatus
ENSPREG00000021082	CRACR2A	97	33.475	ENSMAMG00000005360	cracr2b	95	33.475	Mastacembelus_armatus
ENSPREG00000021082	CRACR2A	92	63.846	ENSMAMG00000011388	cracr2ab	88	63.291	Mastacembelus_armatus
ENSPREG00000021082	CRACR2A	100	71.664	ENSMZEG00005016858	CRACR2A	100	71.664	Maylandia_zebra
ENSPREG00000021082	CRACR2A	95	54.215	ENSMZEG00005005975	cracr2ab	97	54.215	Maylandia_zebra
ENSPREG00000021082	CRACR2A	94	52.074	ENSMAUG00000010171	-	94	52.074	Mesocricetus_auratus
ENSPREG00000021082	CRACR2A	98	52.387	ENSMICG00000012907	CRACR2A	98	52.387	Microcebus_murinus
ENSPREG00000021082	CRACR2A	93	52.933	ENSMOCG00000016052	-	93	52.933	Microtus_ochrogaster
ENSPREG00000021082	CRACR2A	100	67.711	ENSMMOG00000019358	CRACR2A	100	67.439	Mola_mola
ENSPREG00000021082	CRACR2A	95	57.143	ENSMMOG00000001929	cracr2ab	97	57.368	Mola_mola
ENSPREG00000021082	CRACR2A	99	35.007	ENSMALG00000006310	cracr2b	97	35.007	Monopterus_albus
ENSPREG00000021082	CRACR2A	100	69.399	ENSMALG00000021818	CRACR2A	100	69.441	Monopterus_albus
ENSPREG00000021082	CRACR2A	86	62.428	ENSMALG00000012964	cracr2ab	96	50.000	Monopterus_albus
ENSPREG00000021082	CRACR2A	94	53.152	ENSMUSG00000061414	Cracr2a	94	53.143	Mus_musculus
ENSPREG00000021082	CRACR2A	93	53.478	ENSMPUG00000009336	CRACR2A	95	53.478	Mustela_putorius_furo
ENSPREG00000021082	CRACR2A	94	54.011	ENSNGAG00000014625	-	94	54.011	Nannospalax_galili
ENSPREG00000021082	CRACR2A	95	55.800	ENSNBRG00000005924	cracr2ab	97	55.800	Neolamprologus_brichardi
ENSPREG00000021082	CRACR2A	86	72.751	ENSNBRG00000015888	CRACR2A	100	72.751	Neolamprologus_brichardi
ENSPREG00000021082	CRACR2A	96	51.836	ENSNLEG00000003001	CRACR2A	95	52.401	Nomascus_leucogenys
ENSPREG00000021082	CRACR2A	94	51.498	ENSODEG00000002836	CRACR2A	95	51.641	Octodon_degus
ENSPREG00000021082	CRACR2A	94	52.586	ENSOCUG00000005317	CRACR2A	94	52.586	Oryctolagus_cuniculus
ENSPREG00000021082	CRACR2A	100	66.392	ENSORLG00000025905	CRACR2A	100	66.255	Oryzias_latipes
ENSPREG00000021082	CRACR2A	96	48.497	ENSORLG00000014854	cracr2ab	99	48.497	Oryzias_latipes
ENSPREG00000021082	CRACR2A	100	67.129	ENSORLG00020010617	CRACR2A	100	66.990	Oryzias_latipes_hni
ENSPREG00000021082	CRACR2A	100	67.220	ENSORLG00015001413	CRACR2A	100	67.082	Oryzias_latipes_hsok
ENSPREG00000021082	CRACR2A	96	47.700	ENSORLG00015013042	cracr2ab	99	47.700	Oryzias_latipes_hsok
ENSPREG00000021082	CRACR2A	95	54.240	ENSOMEG00000007213	cracr2ab	98	54.240	Oryzias_melastigma
ENSPREG00000021082	CRACR2A	96	53.173	ENSOGAG00000001502	CRACR2A	99	53.173	Otolemur_garnettii
ENSPREG00000021082	CRACR2A	94	54.155	ENSOARG00000010465	CRACR2A	94	54.441	Ovis_aries
ENSPREG00000021082	CRACR2A	74	49.738	ENSOARG00000013673	-	74	49.738	Ovis_aries
ENSPREG00000021082	CRACR2A	96	52.186	ENSPPAG00000026014	CRACR2A	95	53.173	Pan_paniscus
ENSPREG00000021082	CRACR2A	94	53.026	ENSPPRG00000006970	CRACR2A	94	53.314	Panthera_pardus
ENSPREG00000021082	CRACR2A	96	51.893	ENSPTRG00000004545	CRACR2A	95	53.032	Pan_troglodytes
ENSPREG00000021082	CRACR2A	94	53.050	ENSPANG00000025412	CRACR2A	93	53.945	Papio_anubis
ENSPREG00000021082	CRACR2A	96	58.392	ENSPKIG00000019679	-	96	58.392	Paramormyrops_kingsleyae
ENSPREG00000021082	CRACR2A	97	59.607	ENSPKIG00000001449	-	98	59.607	Paramormyrops_kingsleyae
ENSPREG00000021082	CRACR2A	90	63.482	ENSPMGG00000005501	-	90	63.482	Periophthalmus_magnuspinnatus
ENSPREG00000021082	CRACR2A	95	57.849	ENSPMGG00000009818	cracr2ab	97	58.285	Periophthalmus_magnuspinnatus
ENSPREG00000021082	CRACR2A	56	55.072	ENSPCIG00000001087	-	99	55.072	Phascolarctos_cinereus
ENSPREG00000021082	CRACR2A	93	34.200	ENSPFOG00000010947	cracr2b	81	47.895	Poecilia_formosa
ENSPREG00000021082	CRACR2A	94	56.547	ENSPFOG00000005266	cracr2ab	99	56.835	Poecilia_formosa
ENSPREG00000021082	CRACR2A	75	47.895	ENSPLAG00000003773	cracr2b	81	47.895	Poecilia_latipinna
ENSPREG00000021082	CRACR2A	100	96.528	ENSPLAG00000001077	-	100	96.528	Poecilia_latipinna
ENSPREG00000021082	CRACR2A	96	55.696	ENSPLAG00000003905	cracr2ab	98	56.366	Poecilia_latipinna
ENSPREG00000021082	CRACR2A	95	56.579	ENSPMEG00000007308	cracr2ab	98	56.871	Poecilia_mexicana
ENSPREG00000021082	CRACR2A	100	96.250	ENSPMEG00000005485	-	100	96.250	Poecilia_mexicana
ENSPREG00000021082	CRACR2A	75	48.421	ENSPMEG00000020754	cracr2b	81	48.421	Poecilia_mexicana
ENSPREG00000021082	CRACR2A	96	53.090	ENSPCOG00000020329	CRACR2A	95	53.511	Propithecus_coquereli
ENSPREG00000021082	CRACR2A	95	55.764	ENSPNYG00000000606	cracr2ab	97	55.764	Pundamilia_nyererei
ENSPREG00000021082	CRACR2A	95	69.198	ENSPNYG00000020714	CRACR2A	98	69.198	Pundamilia_nyererei
ENSPREG00000021082	CRACR2A	96	55.780	ENSPNAG00000029103	cracr2ab	96	55.780	Pygocentrus_nattereri
ENSPREG00000021082	CRACR2A	100	60.840	ENSPNAG00000020348	CRACR2A	100	61.081	Pygocentrus_nattereri
ENSPREG00000021082	CRACR2A	96	53.156	ENSRBIG00000044914	CRACR2A	95	53.576	Rhinopithecus_bieti
ENSPREG00000021082	CRACR2A	78	53.833	ENSRROG00000028125	CRACR2A	96	53.833	Rhinopithecus_roxellana
ENSPREG00000021082	CRACR2A	96	53.586	ENSSBOG00000024718	CRACR2A	95	54.482	Saimiri_boliviensis_boliviensis
ENSPREG00000021082	CRACR2A	95	62.011	ENSSFOG00015008628	-	99	62.412	Scleropages_formosus
ENSPREG00000021082	CRACR2A	94	63.097	ENSSFOG00015004037	cracr2ab	97	63.689	Scleropages_formosus
ENSPREG00000021082	CRACR2A	94	75.180	ENSSMAG00000011200	CRACR2A	100	72.666	Scophthalmus_maximus
ENSPREG00000021082	CRACR2A	88	66.201	ENSSMAG00000001304	cracr2ab	87	62.341	Scophthalmus_maximus
ENSPREG00000021082	CRACR2A	85	46.119	ENSSDUG00000013283	cracr2b	93	46.119	Seriola_dumerili
ENSPREG00000021082	CRACR2A	100	73.735	ENSSDUG00000000911	CRACR2A	100	73.735	Seriola_dumerili
ENSPREG00000021082	CRACR2A	85	46.364	ENSSLDG00000022995	cracr2b	93	46.364	Seriola_lalandi_dorsalis
ENSPREG00000021082	CRACR2A	95	54.532	ENSSLDG00000023518	cracr2ab	95	54.532	Seriola_lalandi_dorsalis
ENSPREG00000021082	CRACR2A	100	74.008	ENSSLDG00000010742	CRACR2A	100	74.008	Seriola_lalandi_dorsalis
ENSPREG00000021082	CRACR2A	94	54.191	ENSSPUG00000017268	CRACR2A	93	54.665	Sphenodon_punctatus
ENSPREG00000021082	CRACR2A	85	57.368	ENSSPUG00000010462	-	62	57.368	Sphenodon_punctatus
ENSPREG00000021082	CRACR2A	95	54.908	ENSSPAG00000022330	cracr2ab	99	54.908	Stegastes_partitus
ENSPREG00000021082	CRACR2A	100	73.806	ENSSPAG00000012064	CRACR2A	100	73.806	Stegastes_partitus
ENSPREG00000021082	CRACR2A	94	53.546	ENSSSCG00000000732	CRACR2A	99	53.546	Sus_scrofa
ENSPREG00000021082	CRACR2A	98	68.732	ENSTRUG00000023284	CRACR2A	96	68.583	Takifugu_rubripes
ENSPREG00000021082	CRACR2A	93	53.623	ENSUAMG00000021584	CRACR2A	95	53.623	Ursus_americanus
ENSPREG00000021082	CRACR2A	93	53.478	ENSUMAG00000014780	CRACR2A	95	53.478	Ursus_maritimus
ENSPREG00000021082	CRACR2A	94	53.458	ENSVVUG00000018097	CRACR2A	94	53.890	Vulpes_vulpes
ENSPREG00000021082	CRACR2A	97	88.905	ENSXCOG00000007494	CRACR2A	99	88.905	Xiphophorus_couchianus
ENSPREG00000021082	CRACR2A	92	54.674	ENSXCOG00000013604	cracr2ab	99	54.674	Xiphophorus_couchianus
ENSPREG00000021082	CRACR2A	96	55.971	ENSXMAG00000011202	cracr2ab	99	55.971	Xiphophorus_maculatus
ENSPREG00000021082	CRACR2A	100	94.167	ENSXMAG00000021050	CRACR2A	100	94.167	Xiphophorus_maculatus