EuRBPDB

Description

RBDs

Transcripts

Gene Model

PPI

Paralogs

Orthologs

Description

Ensembl ID: ENSSTOG00000003553 (Gene tree)
Gene ID: 101956947
Gene Symbol: LARP4
Alias: PP13296
Full Name: La ribonucleoprotein domain family member 4
Gene Type: protein_coding
Species: Ictidomys_tridecemlineatus
Status: putative
Strand: Plus strand
Length: 72029 bases
Position: chrJH393322.1:8023891-8095919
Accession: 101956947
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
ENSSTOP00000026799	La	PF05383.17	8.1e-18	1	1
ENSSTOP00000003190	La	PF05383.17	8.1e-18	1	1

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
ENSSTOT00000035810	LARP4-201	7284	-	ENSSTOP00000026799	723 (aa)	-	A0A287CZX9
ENSSTOT00000003561	LARP4-202	2882	XM_005324940	ENSSTOP00000003190	724 (aa)	XP_005324997	I3M2I1

Gene Model

Click here to download ENSSTOG00000003553's gene model file

Protein-Protein Interaction (PPI)

Clik here to download ENSSTOG00000003553's network

Paralogs

Ensembl ID	Gene Symbol	Coverage	Identiy	Paralog	Gene Symbol	Coverage	Identiy
ENSSTOG00000003553	LARP4	93	50.390	ENSSTOG00000000587	LARP4B	89	50.390

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
ENSSTOG00000003553	LARP4	100	89.365	ENSG00000161813	LARP4	100	94.595	Homo_sapiens
ENSSTOG00000003553	LARP4	99	46.545	ENSAPOG00000021331	larp4ab	99	46.719	Acanthochromis_polyacanthus
ENSSTOG00000003553	LARP4	99	89.889	ENSAMEG00000001275	LARP4	100	89.889	Ailuropoda_melanoleuca
ENSSTOG00000003553	LARP4	88	63.542	ENSACIG00000001198	larp4aa	94	72.781	Amphilophus_citrinellus
ENSSTOG00000003553	LARP4	99	46.113	ENSAOCG00000006387	larp4ab	99	46.124	Amphiprion_ocellaris
ENSSTOG00000003553	LARP4	51	60.638	ENSAPEG00000015441	larp4ab	98	60.638	Amphiprion_percula
ENSSTOG00000003553	LARP4	99	44.041	ENSATEG00000001018	larp4ab	99	44.228	Anabas_testudineus
ENSSTOG00000003553	LARP4	99	71.390	ENSAPLG00000013661	LARP4	99	70.981	Anas_platyrhynchos
ENSSTOG00000003553	LARP4	100	65.489	ENSACAG00000017133	LARP4	97	65.761	Anolis_carolinensis
ENSSTOG00000003553	LARP4	100	88.372	ENSANAG00000029342	LARP4	97	92.513	Aotus_nancymaae
ENSSTOG00000003553	LARP4	52	63.874	ENSACLG00000002575	larp4aa	95	48.790	Astatotilapia_calliptera
ENSSTOG00000003553	LARP4	51	56.545	ENSACLG00000000777	larp4ab	99	56.283	Astatotilapia_calliptera
ENSSTOG00000003553	LARP4	99	50.514	ENSAMXG00000036493	larp4aa	94	51.421	Astyanax_mexicanus
ENSSTOG00000003553	LARP4	99	46.013	ENSAMXG00000001515	larp4ab	95	46.623	Astyanax_mexicanus
ENSSTOG00000003553	LARP4	100	87.891	ENSBTAG00000017164	LARP4	100	87.891	Bos_taurus
ENSSTOG00000003553	LARP4	100	88.966	ENSCJAG00000021131	LARP4	97	93.048	Callithrix_jacchus
ENSSTOG00000003553	LARP4	100	91.148	ENSCAFG00000008174	LARP4	100	91.148	Canis_familiaris
ENSSTOG00000003553	LARP4	100	90.646	ENSCAFG00020014554	LARP4	100	90.720	Canis_lupus_dingo
ENSSTOG00000003553	LARP4	100	87.483	ENSCHIG00000025965	LARP4	100	87.483	Capra_hircus
ENSSTOG00000003553	LARP4	100	89.241	ENSTSYG00000004342	LARP4	98	92.513	Carlito_syrichta
ENSSTOG00000003553	LARP4	99	68.443	ENSCAPG00000012942	LARP4	100	67.882	Cavia_aperea
ENSSTOG00000003553	LARP4	100	82.735	ENSCPOG00000020519	LARP4	100	82.735	Cavia_porcellus
ENSSTOG00000003553	LARP4	100	88.674	ENSCCAG00000024439	LARP4	97	92.513	Cebus_capucinus
ENSSTOG00000003553	LARP4	99	87.900	ENSCATG00000044528	LARP4	100	87.900	Cercocebus_atys
ENSSTOG00000003553	LARP4	100	87.967	ENSCLAG00000001829	LARP4	100	87.967	Chinchilla_lanigera
ENSSTOG00000003553	LARP4	59	91.429	ENSCHOG00000010490	-	58	91.429	Choloepus_hoffmanni
ENSSTOG00000003553	LARP4	100	72.343	ENSCPBG00000019407	LARP4	91	72.381	Chrysemys_picta_bellii
ENSSTOG00000003553	LARP4	100	80.387	ENSCANG00000033097	-	100	84.791	Colobus_angolensis_palliatus
ENSSTOG00000003553	LARP4	100	89.931	ENSCANG00000029159	-	97	94.385	Colobus_angolensis_palliatus
ENSSTOG00000003553	LARP4	100	87.155	ENSCGRG00001010818	Larp4	100	87.155	Cricetulus_griseus_chok1gshd
ENSSTOG00000003553	LARP4	100	86.999	ENSCGRG00000009382	Larp4	100	87.047	Cricetulus_griseus_crigri
ENSSTOG00000003553	LARP4	92	64.815	ENSCSEG00000001963	larp4aa	81	64.815	Cynoglossus_semilaevis
ENSSTOG00000003553	LARP4	59	55.405	ENSCSEG00000003240	larp4ab	94	44.993	Cynoglossus_semilaevis
ENSSTOG00000003553	LARP4	89	63.542	ENSCVAG00000005488	larp4aa	83	63.542	Cyprinodon_variegatus
ENSSTOG00000003553	LARP4	99	48.000	ENSDARG00000078401	larp4aa	95	48.267	Danio_rerio
ENSSTOG00000003553	LARP4	100	48.312	ENSDARG00000074979	larp4ab	85	80.132	Danio_rerio
ENSSTOG00000003553	LARP4	100	87.724	ENSDNOG00000038525	LARP4	100	87.724	Dasypus_novemcinctus
ENSSTOG00000003553	LARP4	100	88.122	ENSDORG00000004013	Larp4	100	88.122	Dipodomys_ordii
ENSSTOG00000003553	LARP4	73	80.377	ENSETEG00000011683	LARP4	73	80.377	Echinops_telfairi
ENSSTOG00000003553	LARP4	79	39.604	ENSEBUG00000010394	LARP4	87	39.965	Eptatretus_burgeri
ENSSTOG00000003553	LARP4	70	42.572	ENSEBUG00000004426	LARP4	62	52.033	Eptatretus_burgeri
ENSSTOG00000003553	LARP4	99	88.658	ENSEASG00005004356	LARP4	100	92.241	Equus_asinus_asinus
ENSSTOG00000003553	LARP4	100	91.286	ENSECAG00000012479	LARP4	100	91.692	Equus_caballus
ENSSTOG00000003553	LARP4	76	96.581	ENSEEUG00000000316	LARP4	76	96.581	Erinaceus_europaeus
ENSSTOG00000003553	LARP4	88	61.111	ENSELUG00000000395	larp4aa	84	61.111	Esox_lucius
ENSSTOG00000003553	LARP4	99	46.834	ENSELUG00000018650	larp4ab	86	74.894	Esox_lucius
ENSSTOG00000003553	LARP4	100	91.310	ENSFCAG00000031897	LARP4	100	91.310	Felis_catus
ENSSTOG00000003553	LARP4	86	66.770	ENSFALG00000002114	LARP4	98	69.243	Ficedula_albicollis
ENSSTOG00000003553	LARP4	100	87.569	ENSFDAG00000006406	-	100	87.569	Fukomys_damarensis
ENSSTOG00000003553	LARP4	99	43.634	ENSFHEG00000013854	larp4ab	99	43.293	Fundulus_heteroclitus
ENSSTOG00000003553	LARP4	99	49.606	ENSGMOG00000008246	larp4aa	95	49.344	Gadus_morhua
ENSSTOG00000003553	LARP4	95	70.000	ENSGALG00000042122	LARP4	93	69.571	Gallus_gallus
ENSSTOG00000003553	LARP4	99	44.137	ENSGAFG00000021777	larp4ab	99	43.610	Gambusia_affinis
ENSSTOG00000003553	LARP4	99	50.721	ENSGACG00000008381	larp4aa	99	65.517	Gasterosteus_aculeatus
ENSSTOG00000003553	LARP4	99	45.910	ENSGACG00000000680	larp4ab	99	45.778	Gasterosteus_aculeatus
ENSSTOG00000003553	LARP4	100	71.117	ENSGAGG00000006969	LARP4	100	71.156	Gopherus_agassizii
ENSSTOG00000003553	LARP4	100	84.924	ENSGGOG00000015230	LARP4	99	93.531	Gorilla_gorilla
ENSSTOG00000003553	LARP4	52	63.874	ENSHBUG00000023074	larp4aa	95	48.662	Haplochromis_burtoni
ENSSTOG00000003553	LARP4	91	86.254	ENSHGLG00000005513	-	100	86.254	Heterocephalus_glaber_female
ENSSTOG00000003553	LARP4	86	81.583	ENSHGLG00000013875	-	100	80.614	Heterocephalus_glaber_female
ENSSTOG00000003553	LARP4	62	75.055	ENSHGLG00000010979	-	93	76.871	Heterocephalus_glaber_female
ENSSTOG00000003553	LARP4	100	86.538	ENSHGLG00100016984	-	100	86.538	Heterocephalus_glaber_male
ENSSTOG00000003553	LARP4	100	81.604	ENSHGLG00100011678	-	100	81.077	Heterocephalus_glaber_male
ENSSTOG00000003553	LARP4	62	75.055	ENSHGLG00100002832	-	93	76.871	Heterocephalus_glaber_male
ENSSTOG00000003553	LARP4	99	41.689	ENSHCOG00000020857	larp4ab	99	41.557	Hippocampus_comes
ENSSTOG00000003553	LARP4	99	48.249	ENSHCOG00000015951	larp4aa	95	48.964	Hippocampus_comes
ENSSTOG00000003553	LARP4	99	48.395	ENSIPUG00000007514	larp4aa	95	48.780	Ictalurus_punctatus
ENSSTOG00000003553	LARP4	100	47.867	ENSIPUG00000021659	larp4ab	100	47.145	Ictalurus_punctatus
ENSSTOG00000003553	LARP4	52	69.474	ENSJJAG00000014649	-	95	89.048	Jaculus_jaculus
ENSSTOG00000003553	LARP4	99	45.621	ENSKMAG00000018556	larp4ab	99	45.178	Kryptolebias_marmoratus
ENSSTOG00000003553	LARP4	99	44.110	ENSLBEG00000019306	larp4ab	100	43.603	Labrus_bergylta
ENSSTOG00000003553	LARP4	99	60.188	ENSLACG00000019090	LARP4	99	58.713	Latimeria_chalumnae
ENSSTOG00000003553	LARP4	99	53.706	ENSLOCG00000007113	larp4ab	95	52.995	Lepisosteus_oculatus
ENSSTOG00000003553	LARP4	100	88.536	ENSLAFG00000005160	LARP4	100	88.536	Loxodonta_africana
ENSSTOG00000003553	LARP4	99	90.014	ENSMFAG00000043681	LARP4	97	94.385	Macaca_fascicularis
ENSSTOG00000003553	LARP4	100	89.330	ENSMMUG00000008172	LARP4	97	94.385	Macaca_mulatta
ENSSTOG00000003553	LARP4	87	72.277	ENSMNEG00000037799	-	99	75.806	Macaca_nemestrina
ENSSTOG00000003553	LARP4	93	90.448	ENSMNEG00000034602	LARP4	100	88.702	Macaca_nemestrina
ENSSTOG00000003553	LARP4	100	90.345	ENSMLEG00000039661	LARP4	97	94.920	Mandrillus_leucophaeus
ENSSTOG00000003553	LARP4	99	44.401	ENSMAMG00000002616	larp4ab	99	44.603	Mastacembelus_armatus
ENSSTOG00000003553	LARP4	99	48.214	ENSMZEG00005001813	larp4aa	95	48.790	Maylandia_zebra
ENSSTOG00000003553	LARP4	51	56.545	ENSMZEG00005007771	larp4ab	99	56.283	Maylandia_zebra
ENSSTOG00000003553	LARP4	95	69.286	ENSMGAG00000010598	LARP4	93	69.088	Meleagris_gallopavo
ENSSTOG00000003553	LARP4	99	84.262	ENSMICG00000030156	-	99	84.262	Microcebus_murinus
ENSSTOG00000003553	LARP4	87	90.016	ENSMICG00000012388	-	99	90.016	Microcebus_murinus
ENSSTOG00000003553	LARP4	100	85.773	ENSMOCG00000020635	Larp4	100	85.773	Microtus_ochrogaster
ENSSTOG00000003553	LARP4	100	42.989	ENSMMOG00000021861	larp4ab	94	54.098	Mola_mola
ENSSTOG00000003553	LARP4	95	76.224	MGP_CAROLIEiJ_G0020281	Larp4	85	91.212	Mus_caroli
ENSSTOG00000003553	LARP4	100	85.912	ENSMUSG00000023025	Larp4	100	86.103	Mus_musculus
ENSSTOG00000003553	LARP4	100	86.464	MGP_PahariEiJ_G0020284	Larp4	100	86.464	Mus_pahari
ENSSTOG00000003553	LARP4	99	90.278	ENSMPUG00000008706	LARP4	96	90.278	Mustela_putorius_furo
ENSSTOG00000003553	LARP4	100	86.621	ENSMLUG00000012513	-	100	86.520	Myotis_lucifugus
ENSSTOG00000003553	LARP4	100	86.326	ENSNGAG00000001577	-	100	86.326	Nannospalax_galili
ENSSTOG00000003553	LARP4	51	56.117	ENSNBRG00000015675	larp4ab	99	56.117	Neolamprologus_brichardi
ENSSTOG00000003553	LARP4	99	47.170	ENSNBRG00000011292	larp4aa	95	48.593	Neolamprologus_brichardi
ENSSTOG00000003553	LARP4	100	90.055	ENSNLEG00000017798	LARP4	97	93.583	Nomascus_leucogenys
ENSSTOG00000003553	LARP4	67	74.583	ENSOPRG00000004592	-	80	74.583	Ochotona_princeps
ENSSTOG00000003553	LARP4	99	84.262	ENSODEG00000009770	LARP4	100	84.262	Octodon_degus
ENSSTOG00000003553	LARP4	100	43.512	ENSONIG00000019124	larp4ab	100	42.839	Oreochromis_niloticus
ENSSTOG00000003553	LARP4	100	49.673	ENSONIG00000011895	larp4aa	99	49.935	Oreochromis_niloticus
ENSSTOG00000003553	LARP4	73	71.852	ENSOANG00000004373	LARP4	97	71.852	Ornithorhynchus_anatinus
ENSSTOG00000003553	LARP4	100	87.172	ENSOCUG00000021417	-	100	87.172	Oryctolagus_cuniculus
ENSSTOG00000003553	LARP4	100	87.862	ENSOCUG00000026872	-	99	86.345	Oryctolagus_cuniculus
ENSSTOG00000003553	LARP4	99	50.783	ENSORLG00000006758	larp4aa	95	49.870	Oryzias_latipes
ENSSTOG00000003553	LARP4	99	50.261	ENSORLG00020010258	larp4aa	95	49.349	Oryzias_latipes_hni
ENSSTOG00000003553	LARP4	99	50.914	ENSORLG00015002292	larp4aa	95	49.294	Oryzias_latipes_hsok
ENSSTOG00000003553	LARP4	99	50.199	ENSOMEG00000019749	larp4aa	97	66.575	Oryzias_melastigma
ENSSTOG00000003553	LARP4	71	88.953	ENSOGAG00000001415	LARP4	99	88.953	Otolemur_garnettii
ENSSTOG00000003553	LARP4	100	87.176	ENSOARG00000017702	LARP4	99	87.176	Ovis_aries
ENSSTOG00000003553	LARP4	90	91.837	ENSPPAG00000035100	LARP4	97	93.048	Pan_paniscus
ENSSTOG00000003553	LARP4	62	72.627	ENSPPAG00000039122	-	99	71.429	Pan_paniscus
ENSSTOG00000003553	LARP4	100	91.172	ENSPPRG00000004522	LARP4	100	91.172	Panthera_pardus
ENSSTOG00000003553	LARP4	100	91.172	ENSPTIG00000015617	LARP4	100	91.172	Panthera_tigris_altaica
ENSSTOG00000003553	LARP4	100	89.379	ENSPTRG00000004935	LARP4	97	93.316	Pan_troglodytes
ENSSTOG00000003553	LARP4	62	72.406	ENSPTRG00000045907	-	98	79.404	Pan_troglodytes
ENSSTOG00000003553	LARP4	100	89.603	ENSPANG00000022964	LARP4	97	94.385	Papio_anubis
ENSSTOG00000003553	LARP4	100	50.377	ENSPKIG00000005950	larp4ab	100	49.499	Paramormyrops_kingsleyae
ENSSTOG00000003553	LARP4	99	69.505	ENSPSIG00000015867	LARP4	99	69.505	Pelodiscus_sinensis
ENSSTOG00000003553	LARP4	50	54.497	ENSPMGG00000002719	larp4ab	97	55.026	Periophthalmus_magnuspinnatus
ENSSTOG00000003553	LARP4	99	80.690	ENSPEMG00000018433	Larp4	94	86.526	Peromyscus_maniculatus_bairdii
ENSSTOG00000003553	LARP4	100	76.054	ENSPCIG00000000384	LARP4	100	75.782	Phascolarctos_cinereus
ENSSTOG00000003553	LARP4	99	44.993	ENSPFOG00000009764	larp4ab	99	44.740	Poecilia_formosa
ENSSTOG00000003553	LARP4	99	49.239	ENSPFOG00000013148	larp4aa	100	48.680	Poecilia_formosa
ENSSTOG00000003553	LARP4	99	49.301	ENSPLAG00000013626	larp4aa	95	49.872	Poecilia_latipinna
ENSSTOG00000003553	LARP4	99	44.533	ENSPLAG00000008502	larp4ab	99	44.282	Poecilia_latipinna
ENSSTOG00000003553	LARP4	99	49.239	ENSPMEG00000014271	larp4aa	95	49.809	Poecilia_mexicana
ENSSTOG00000003553	LARP4	99	44.386	ENSPMEG00000002720	larp4ab	99	44.254	Poecilia_mexicana
ENSSTOG00000003553	LARP4	99	50.063	ENSPREG00000022597	larp4aa	95	49.744	Poecilia_reticulata
ENSSTOG00000003553	LARP4	51	59.043	ENSPREG00000016263	larp4ab	99	57.769	Poecilia_reticulata
ENSSTOG00000003553	LARP4	71	92.054	ENSPPYG00000004512	-	100	92.054	Pongo_abelii
ENSSTOG00000003553	LARP4	93	80.149	ENSPCAG00000003671	LARP4	99	80.149	Procavia_capensis
ENSSTOG00000003553	LARP4	88	91.957	ENSPCOG00000026291	LARP4	98	91.979	Propithecus_coquereli
ENSSTOG00000003553	LARP4	93	79.851	ENSPVAG00000016840	LARP4	91	79.851	Pteropus_vampyrus
ENSSTOG00000003553	LARP4	52	63.874	ENSPNYG00000022265	larp4aa	95	48.790	Pundamilia_nyererei
ENSSTOG00000003553	LARP4	100	47.516	ENSPNAG00000014478	larp4ab	100	48.849	Pygocentrus_nattereri
ENSSTOG00000003553	LARP4	96	50.134	ENSPNAG00000024770	larp4aa	94	50.804	Pygocentrus_nattereri
ENSSTOG00000003553	LARP4	88	93.801	ENSRBIG00000038857	-	99	93.197	Rhinopithecus_bieti
ENSSTOG00000003553	LARP4	89	81.102	ENSRBIG00000036890	-	100	81.102	Rhinopithecus_bieti
ENSSTOG00000003553	LARP4	100	89.163	ENSRROG00000042764	-	99	93.424	Rhinopithecus_roxellana
ENSSTOG00000003553	LARP4	100	80.110	ENSRROG00000025487	-	100	81.365	Rhinopithecus_roxellana
ENSSTOG00000003553	LARP4	100	88.552	ENSSBOG00000026650	LARP4	100	89.466	Saimiri_boliviensis_boliviensis
ENSSTOG00000003553	LARP4	100	51.706	ENSSFOG00015003868	larp4aa	100	50.980	Scleropages_formosus
ENSSTOG00000003553	LARP4	100	49.738	ENSSFOG00015014096	larp4ab	100	50.000	Scleropages_formosus
ENSSTOG00000003553	LARP4	99	46.354	ENSSMAG00000002838	larp4ab	99	47.204	Scophthalmus_maximus
ENSSTOG00000003553	LARP4	99	46.572	ENSSDUG00000006655	larp4ab	99	46.521	Seriola_dumerili
ENSSTOG00000003553	LARP4	99	46.563	ENSSLDG00000001791	larp4ab	99	46.521	Seriola_lalandi_dorsalis
ENSSTOG00000003553	LARP4	99	81.793	ENSSARG00000012469	LARP4	100	81.793	Sorex_araneus
ENSSTOG00000003553	LARP4	95	69.318	ENSSPUG00000016552	LARP4	93	69.602	Sphenodon_punctatus
ENSSTOG00000003553	LARP4	99	49.281	ENSSPAG00000007222	larp4aa	98	63.172	Stegastes_partitus
ENSSTOG00000003553	LARP4	99	46.466	ENSSPAG00000002172	larp4ab	99	45.241	Stegastes_partitus
ENSSTOG00000003553	LARP4	100	89.256	ENSSSCG00000028516	LARP4	97	92.876	Sus_scrofa
ENSSTOG00000003553	LARP4	100	68.033	ENSTGUG00000003479	LARP4	100	68.759	Taeniopygia_guttata
ENSSTOG00000003553	LARP4	50	60.773	ENSTRUG00000015939	larp4aa	100	60.773	Takifugu_rubripes
ENSSTOG00000003553	LARP4	99	45.321	ENSTNIG00000014571	larp4ab	54	59.791	Tetraodon_nigroviridis
ENSSTOG00000003553	LARP4	99	48.553	ENSTNIG00000015109	larp4aa	50	62.078	Tetraodon_nigroviridis
ENSSTOG00000003553	LARP4	99	78.177	ENSTBEG00000005167	LARP4	100	78.177	Tupaia_belangeri
ENSSTOG00000003553	LARP4	100	87.260	ENSTTRG00000003764	LARP4	99	87.862	Tursiops_truncatus
ENSSTOG00000003553	LARP4	72	90.366	ENSUAMG00000013764	LARP4	99	92.744	Ursus_americanus
ENSSTOG00000003553	LARP4	100	89.793	ENSUMAG00000008473	LARP4	100	89.793	Ursus_maritimus
ENSSTOG00000003553	LARP4	100	87.211	ENSVPAG00000010136	LARP4	100	86.821	Vicugna_pacos
ENSSTOG00000003553	LARP4	100	90.608	ENSVVUG00000024717	LARP4	97	94.709	Vulpes_vulpes
ENSSTOG00000003553	LARP4	93	60.405	ENSXETG00000007283	larp4	100	60.405	Xenopus_tropicalis
ENSSTOG00000003553	LARP4	51	58.158	ENSXCOG00000019635	larp4ab	97	58.158	Xiphophorus_couchianus
ENSSTOG00000003553	LARP4	99	48.473	ENSXCOG00000006387	larp4aa	95	47.954	Xiphophorus_couchianus
ENSSTOG00000003553	LARP4	99	45.443	ENSXMAG00000002156	larp4ab	99	44.914	Xiphophorus_maculatus
ENSSTOG00000003553	LARP4	99	49.564	ENSXMAG00000010519	larp4aa	95	50.384	Xiphophorus_maculatus