EuRBPDB

Description

RBDs

Transcripts

Gene Model

PPI

Paralogs

Orthologs

Description

Ensembl ID: ENSUAMG00000001610 (Gene tree)
Gene ID: -
Gene Symbol: -
Alias: IMP-2
Full Name: -
Gene Type: protein_coding
Species: Ursus_americanus
Status: putative
Strand: Plus strand
Length: 26467 bases
Position: chrLZNR01014265.1:10705-37171
Accession: -
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
ENSUAMP00000002413	KH_1	PF00013.29	2.5e-42	1	3
ENSUAMP00000002413	KH_1	PF00013.29	2.5e-42	2	3
ENSUAMP00000002413	KH_1	PF00013.29	2.5e-42	3	3
ENSUAMP00000002404	KH_1	PF00013.29	2.6e-42	1	3
ENSUAMP00000002404	KH_1	PF00013.29	2.6e-42	2	3
ENSUAMP00000002404	KH_1	PF00013.29	2.6e-42	3	3
ENSUAMP00000002363	KH_1	PF00013.29	4e-42	1	3
ENSUAMP00000002363	KH_1	PF00013.29	4e-42	2	3
ENSUAMP00000002363	KH_1	PF00013.29	4e-42	3	3
ENSUAMP00000002417	KH_1	PF00013.29	4e-42	1	3
ENSUAMP00000002417	KH_1	PF00013.29	4e-42	2	3
ENSUAMP00000002417	KH_1	PF00013.29	4e-42	3	3
ENSUAMP00000002292	KH_1	PF00013.29	4.5e-42	1	3
ENSUAMP00000002292	KH_1	PF00013.29	4.5e-42	2	3
ENSUAMP00000002292	KH_1	PF00013.29	4.5e-42	3	3

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
ENSUAMT00000002755	-	1053	-	ENSUAMP00000002413	350 (aa)	-	-
ENSUAMT00000002744	-	1065	-	ENSUAMP00000002404	354 (aa)	-	-
ENSUAMT00000002759	-	1176	-	ENSUAMP00000002417	391 (aa)	-	-
ENSUAMT00000002698	-	1179	-	ENSUAMP00000002363	392 (aa)	-	-
ENSUAMT00000002620	-	1212	-	ENSUAMP00000002292	403 (aa)	-	-

Gene Model

Click here to download ENSUAMG00000001610's gene model file

Protein-Protein Interaction (PPI)

Clik here to download ENSUAMG00000001610's network

Paralogs

Ensembl ID	Gene Symbol	Coverage	Identiy	Paralog	Gene Symbol	Coverage	Identiy
ENSUAMG00000001610	-	94	67.378	ENSUAMG00000023269	IGF2BP1	68	68.404
ENSUAMG00000001610	-	81	36.364	ENSUAMG00000006491	PCBP3	96	37.037
ENSUAMG00000001610	-	92	68.987	ENSUAMG00000021770	-	78	69.697

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
ENSUAMG00000001610	-	94	98.476	ENSG00000073792	IGF2BP2	75	98.476	Homo_sapiens
ENSUAMG00000001610	-	92	77.676	ENSAPOG00000004098	igf2bp2a	66	75.884	Acanthochromis_polyacanthus
ENSUAMG00000001610	-	94	99.695	ENSAMEG00000016763	IGF2BP2	67	99.695	Ailuropoda_melanoleuca
ENSUAMG00000001610	-	93	75.684	ENSACIG00000001308	igf2bp2b	66	74.772	Amphilophus_citrinellus
ENSUAMG00000001610	-	92	75.776	ENSAOCG00000016871	igf2bp2b	66	74.510	Amphiprion_ocellaris
ENSUAMG00000001610	-	94	77.177	ENSAPEG00000016037	igf2bp2a	66	75.884	Amphiprion_percula
ENSUAMG00000001610	-	92	81.040	ENSATEG00000020389	igf2bp2b	75	80.122	Anabas_testudineus
ENSUAMG00000001610	-	94	83.082	ENSACAG00000007995	IGF2BP2	55	83.082	Anolis_carolinensis
ENSUAMG00000001610	-	94	98.171	ENSANAG00000031810	IGF2BP2	86	98.171	Aotus_nancymaae
ENSUAMG00000001610	-	92	80.428	ENSACLG00000015512	igf2bp2a	75	78.899	Astatotilapia_calliptera
ENSUAMG00000001610	-	71	71.084	ENSAMXG00000029202	-	92	71.548	Astyanax_mexicanus
ENSUAMG00000001610	-	93	75.841	ENSAMXG00000032347	igf2bp2b	71	72.699	Astyanax_mexicanus
ENSUAMG00000001610	-	94	99.390	ENSBTAG00000007666	IGF2BP2	67	99.390	Bos_taurus
ENSUAMG00000001610	-	94	98.171	ENSCJAG00000014731	IGF2BP2	80	98.171	Callithrix_jacchus
ENSUAMG00000001610	-	94	99.695	ENSCAFG00000013297	IGF2BP2	76	99.695	Canis_familiaris
ENSUAMG00000001610	-	94	99.695	ENSCAFG00020005444	IGF2BP2	67	99.695	Canis_lupus_dingo
ENSUAMG00000001610	-	94	99.390	ENSCHIG00000015258	IGF2BP2	77	99.390	Capra_hircus
ENSUAMG00000001610	-	94	98.780	ENSTSYG00000003376	IGF2BP2	77	98.780	Carlito_syrichta
ENSUAMG00000001610	-	94	98.171	ENSCAPG00000003061	IGF2BP2	77	98.171	Cavia_aperea
ENSUAMG00000001610	-	94	98.171	ENSCPOG00000005471	IGF2BP2	75	98.171	Cavia_porcellus
ENSUAMG00000001610	-	94	97.866	ENSCCAG00000034879	IGF2BP2	67	97.866	Cebus_capucinus
ENSUAMG00000001610	-	94	71.341	ENSCATG00000033281	-	59	69.817	Cercocebus_atys
ENSUAMG00000001610	-	94	98.476	ENSCATG00000036407	IGF2BP2	77	98.476	Cercocebus_atys
ENSUAMG00000001610	-	87	50.968	ENSCATG00000035160	-	61	51.475	Cercocebus_atys
ENSUAMG00000001610	-	94	97.256	ENSCLAG00000014808	IGF2BP2	75	97.256	Chinchilla_lanigera
ENSUAMG00000001610	-	94	98.476	ENSCSAG00000014019	IGF2BP2	67	98.476	Chlorocebus_sabaeus
ENSUAMG00000001610	-	57	96.512	ENSCHOG00000009982	-	58	69.079	Choloepus_hoffmanni
ENSUAMG00000001610	-	94	85.106	ENSCPBG00000007370	IGF2BP2	70	85.106	Chrysemys_picta_bellii
ENSUAMG00000001610	-	94	98.476	ENSCANG00000008906	IGF2BP2	77	98.476	Colobus_angolensis_palliatus
ENSUAMG00000001610	-	94	71.646	ENSCANG00000030550	-	60	71.646	Colobus_angolensis_palliatus
ENSUAMG00000001610	-	94	94.817	ENSCGRG00001019401	Igf2bp2	73	94.817	Cricetulus_griseus_chok1gshd
ENSUAMG00000001610	-	94	94.817	ENSCGRG00000016050	Igf2bp2	83	94.817	Cricetulus_griseus_crigri
ENSUAMG00000001610	-	92	77.370	ENSCSEG00000011718	igf2bp2b	69	70.822	Cynoglossus_semilaevis
ENSUAMG00000001610	-	92	80.122	ENSCVAG00000004656	igf2bp2b	66	78.287	Cyprinodon_variegatus
ENSUAMG00000001610	-	93	80.422	ENSDARG00000003421	igf2bp2a	100	78.544	Danio_rerio
ENSUAMG00000001610	-	91	74.062	ENSDARG00000099758	igf2bp2b	85	93.103	Danio_rerio
ENSUAMG00000001610	-	94	99.085	ENSDNOG00000013421	IGF2BP2	76	99.085	Dasypus_novemcinctus
ENSUAMG00000001610	-	94	96.037	ENSDORG00000000886	Igf2bp2	84	96.037	Dipodomys_ordii
ENSUAMG00000001610	-	94	86.890	ENSEASG00005013866	IGF2BP2	66	86.890	Equus_asinus_asinus
ENSUAMG00000001610	-	94	98.476	ENSECAG00000020685	IGF2BP2	67	98.476	Equus_caballus
ENSUAMG00000001610	-	93	82.263	ENSEEUG00000009674	IGF2BP2	61	82.263	Erinaceus_europaeus
ENSUAMG00000001610	-	87	66.330	ENSELUG00000010676	igf2bp2a	86	80.269	Esox_lucius
ENSUAMG00000001610	-	94	99.390	ENSFCAG00000005369	IGF2BP2	75	99.390	Felis_catus
ENSUAMG00000001610	-	94	83.086	ENSFALG00000013346	IGF2BP2	76	83.086	Ficedula_albicollis
ENSUAMG00000001610	-	94	99.085	ENSFDAG00000015039	IGF2BP2	75	99.085	Fukomys_damarensis
ENSUAMG00000001610	-	92	71.429	ENSFHEG00000008263	igf2bp2a	97	73.864	Fundulus_heteroclitus
ENSUAMG00000001610	-	85	78.477	ENSGMOG00000019327	igf2bp2b	63	76.490	Gadus_morhua
ENSUAMG00000001610	-	93	85.976	ENSGALG00000033195	IGF2BP2	65	85.976	Gallus_gallus
ENSUAMG00000001610	-	92	76.855	ENSGAFG00000010504	igf2bp2b	66	77.644	Gambusia_affinis
ENSUAMG00000001610	-	92	78.899	ENSGACG00000005991	igf2bp2b	69	77.982	Gasterosteus_aculeatus
ENSUAMG00000001610	-	94	84.802	ENSGAGG00000008990	IGF2BP2	70	84.802	Gopherus_agassizii
ENSUAMG00000001610	-	94	78.659	ENSGGOG00000024907	-	62	75.305	Gorilla_gorilla
ENSUAMG00000001610	-	94	98.476	ENSGGOG00000015035	IGF2BP2	75	98.476	Gorilla_gorilla
ENSUAMG00000001610	-	94	78.963	ENSGGOG00000038298	-	60	78.963	Gorilla_gorilla
ENSUAMG00000001610	-	92	80.428	ENSHBUG00000003126	igf2bp2a	75	78.899	Haplochromis_burtoni
ENSUAMG00000001610	-	94	97.879	ENSHGLG00000016529	IGF2BP2	89	90.191	Heterocephalus_glaber_female
ENSUAMG00000001610	-	94	98.171	ENSHGLG00100014188	IGF2BP2	75	98.171	Heterocephalus_glaber_male
ENSUAMG00000001610	-	85	76.174	ENSHCOG00000018090	igf2bp2b	100	76.046	Hippocampus_comes
ENSUAMG00000001610	-	94	73.874	ENSIPUG00000022106	Igf2bp2	70	73.574	Ictalurus_punctatus
ENSUAMG00000001610	-	94	97.866	ENSSTOG00000022077	IGF2BP2	67	97.866	Ictidomys_tridecemlineatus
ENSUAMG00000001610	-	94	96.951	ENSJJAG00000015876	Igf2bp2	69	96.951	Jaculus_jaculus
ENSUAMG00000001610	-	92	78.287	ENSKMAG00000012177	igf2bp2b	66	76.758	Kryptolebias_marmoratus
ENSUAMG00000001610	-	91	75.000	ENSLBEG00000003710	IGF2BP2	97	75.984	Labrus_bergylta
ENSUAMG00000001610	-	94	81.928	ENSLACG00000011546	IGF2BP2	100	79.116	Latimeria_chalumnae
ENSUAMG00000001610	-	92	82.407	ENSLOCG00000009564	igf2bp2b	67	79.938	Lepisosteus_oculatus
ENSUAMG00000001610	-	94	99.085	ENSLAFG00000017156	IGF2BP2	78	99.085	Loxodonta_africana
ENSUAMG00000001610	-	78	64.729	ENSMFAG00000002557	-	58	64.729	Macaca_fascicularis
ENSUAMG00000001610	-	94	98.476	ENSMFAG00000039540	IGF2BP2	77	98.476	Macaca_fascicularis
ENSUAMG00000001610	-	94	73.171	ENSMMUG00000043560	-	51	73.171	Macaca_mulatta
ENSUAMG00000001610	-	94	98.476	ENSMMUG00000023416	IGF2BP2	76	98.476	Macaca_mulatta
ENSUAMG00000001610	-	94	50.928	ENSMNEG00000032558	-	70	49.624	Macaca_nemestrina
ENSUAMG00000001610	-	94	98.171	ENSMNEG00000042466	IGF2BP2	77	98.171	Macaca_nemestrina
ENSUAMG00000001610	-	78	67.470	ENSMNEG00000034048	-	61	67.742	Macaca_nemestrina
ENSUAMG00000001610	-	94	98.476	ENSMLEG00000039422	IGF2BP2	77	98.476	Mandrillus_leucophaeus
ENSUAMG00000001610	-	87	54.397	ENSMLEG00000030453	-	63	53.746	Mandrillus_leucophaeus
ENSUAMG00000001610	-	92	81.040	ENSMAMG00000000969	igf2bp2b	69	79.205	Mastacembelus_armatus
ENSUAMG00000001610	-	92	80.122	ENSMZEG00005007777	igf2bp2a	75	78.593	Maylandia_zebra
ENSUAMG00000001610	-	94	85.714	ENSMGAG00000002904	IGF2BP2	80	85.714	Meleagris_gallopavo
ENSUAMG00000001610	-	94	94.207	ENSMAUG00000008302	Igf2bp2	68	94.207	Mesocricetus_auratus
ENSUAMG00000001610	-	94	98.476	ENSMICG00000049453	-	67	98.476	Microcebus_murinus
ENSUAMG00000001610	-	94	88.720	ENSMICG00000037971	-	85	88.720	Microcebus_murinus
ENSUAMG00000001610	-	94	96.037	ENSMOCG00000019099	Igf2bp2	73	96.037	Microtus_ochrogaster
ENSUAMG00000001610	-	92	70.497	ENSMMOG00000003830	igf2bp2b	71	70.497	Mola_mola
ENSUAMG00000001610	-	93	84.639	ENSMODG00000002822	IGF2BP2	81	84.639	Monodelphis_domestica
ENSUAMG00000001610	-	92	80.923	ENSMALG00000016964	igf2bp2b	77	80.000	Monopterus_albus
ENSUAMG00000001610	-	94	95.427	MGP_CAROLIEiJ_G0020601	Igf2bp2	73	95.427	Mus_caroli
ENSUAMG00000001610	-	94	95.732	ENSMUSG00000033581	Igf2bp2	82	95.732	Mus_musculus
ENSUAMG00000001610	-	94	84.146	MGP_PahariEiJ_G0016305	Igf2bp2	71	84.146	Mus_pahari
ENSUAMG00000001610	-	94	95.732	MGP_SPRETEiJ_G0021497	Igf2bp2	73	95.732	Mus_spretus
ENSUAMG00000001610	-	94	99.390	ENSMPUG00000003151	-	65	99.390	Mustela_putorius_furo
ENSUAMG00000001610	-	93	97.866	ENSMLUG00000002631	IGF2BP2	76	97.866	Myotis_lucifugus
ENSUAMG00000001610	-	94	96.341	ENSNGAG00000012680	Igf2bp2	77	96.341	Nannospalax_galili
ENSUAMG00000001610	-	92	80.923	ENSNBRG00000016140	igf2bp2b	69	79.385	Neolamprologus_brichardi
ENSUAMG00000001610	-	94	98.780	ENSNLEG00000006508	IGF2BP2	77	98.780	Nomascus_leucogenys
ENSUAMG00000001610	-	94	76.220	ENSNLEG00000027269	-	61	76.220	Nomascus_leucogenys
ENSUAMG00000001610	-	94	73.171	ENSNLEG00000032080	-	69	67.945	Nomascus_leucogenys
ENSUAMG00000001610	-	94	77.744	ENSOPRG00000002928	IGF2BP2	78	77.744	Ochotona_princeps
ENSUAMG00000001610	-	94	96.049	ENSODEG00000001231	IGF2BP2	75	96.049	Octodon_degus
ENSUAMG00000001610	-	92	79.817	ENSONIG00000009029	igf2bp2b	75	78.287	Oreochromis_niloticus
ENSUAMG00000001610	-	94	83.636	ENSOANG00000004648	IGF2BP2	79	86.834	Ornithorhynchus_anatinus
ENSUAMG00000001610	-	94	98.476	ENSOCUG00000005057	IGF2BP2	67	98.476	Oryctolagus_cuniculus
ENSUAMG00000001610	-	91	81.424	ENSORLG00000016532	igf2bp2b	69	80.495	Oryzias_latipes
ENSUAMG00000001610	-	92	80.183	ENSORLG00020001556	igf2bp2b	68	80.000	Oryzias_latipes_hni
ENSUAMG00000001610	-	92	80.183	ENSORLG00015001778	igf2bp2b	68	80.000	Oryzias_latipes_hsok
ENSUAMG00000001610	-	91	80.000	ENSOMEG00000001908	igf2bp2b	68	79.077	Oryzias_melastigma
ENSUAMG00000001610	-	94	97.281	ENSOGAG00000010577	IGF2BP2	77	97.281	Otolemur_garnettii
ENSUAMG00000001610	-	94	99.085	ENSOARG00000020556	IGF2BP2	67	99.085	Ovis_aries
ENSUAMG00000001610	-	94	80.488	ENSPPAG00000038959	-	74	78.659	Pan_paniscus
ENSUAMG00000001610	-	94	75.610	ENSPPAG00000002120	-	52	75.610	Pan_paniscus
ENSUAMG00000001610	-	94	98.476	ENSPPAG00000034539	IGF2BP2	80	98.476	Pan_paniscus
ENSUAMG00000001610	-	94	75.915	ENSPPAG00000035134	-	52	75.915	Pan_paniscus
ENSUAMG00000001610	-	94	99.390	ENSPPRG00000003987	IGF2BP2	75	99.390	Panthera_pardus
ENSUAMG00000001610	-	94	99.390	ENSPTIG00000013652	IGF2BP2	75	99.390	Panthera_tigris_altaica
ENSUAMG00000001610	-	94	75.305	ENSPTRG00000049696	-	52	75.305	Pan_troglodytes
ENSUAMG00000001610	-	94	98.476	ENSPTRG00000015705	IGF2BP2	80	98.476	Pan_troglodytes
ENSUAMG00000001610	-	94	76.220	ENSPTRG00000046113	-	52	76.220	Pan_troglodytes
ENSUAMG00000001610	-	94	98.476	ENSPANG00000007053	IGF2BP2	75	98.476	Papio_anubis
ENSUAMG00000001610	-	94	73.171	ENSPANG00000033514	-	60	73.171	Papio_anubis
ENSUAMG00000001610	-	92	81.115	ENSPKIG00000009855	igf2bp2b	66	80.495	Paramormyrops_kingsleyae
ENSUAMG00000001610	-	74	87.387	ENSPSIG00000013917	IGF2BP2	60	87.273	Pelodiscus_sinensis
ENSUAMG00000001610	-	87	71.197	ENSPMGG00000011823	igf2bp2a	63	69.281	Periophthalmus_magnuspinnatus
ENSUAMG00000001610	-	66	70.130	ENSPMAG00000005047	igf2bp2b	92	71.493	Petromyzon_marinus
ENSUAMG00000001610	-	87	77.604	ENSPCIG00000023547	-	63	77.604	Phascolarctos_cinereus
ENSUAMG00000001610	-	92	77.381	ENSPFOG00000007221	igf2bp2b	66	78.788	Poecilia_formosa
ENSUAMG00000001610	-	92	71.739	ENSPLAG00000005993	igf2bp2b	64	72.669	Poecilia_latipinna
ENSUAMG00000001610	-	92	86.239	ENSPMEG00000001726	igf2bp2b	61	86.239	Poecilia_mexicana
ENSUAMG00000001610	-	92	87.059	ENSPREG00000009440	igf2bp2b	62	87.059	Poecilia_reticulata
ENSUAMG00000001610	-	87	96.364	ENSPPYG00000014373	-	64	96.364	Pongo_abelii
ENSUAMG00000001610	-	94	78.049	ENSPCAG00000015093	IGF2BP2	76	78.049	Procavia_capensis
ENSUAMG00000001610	-	94	98.476	ENSPCOG00000027031	IGF2BP2	77	98.476	Propithecus_coquereli
ENSUAMG00000001610	-	94	91.768	ENSPVAG00000011494	IGF2BP2	67	91.768	Pteropus_vampyrus
ENSUAMG00000001610	-	92	79.205	ENSPNYG00000001187	igf2bp2b	69	77.676	Pundamilia_nyererei
ENSUAMG00000001610	-	74	71.042	ENSPNAG00000013765	-	91	73.840	Pygocentrus_nattereri
ENSUAMG00000001610	-	93	85.465	ENSPNAG00000011228	igf2bp2b	67	85.465	Pygocentrus_nattereri
ENSUAMG00000001610	-	94	96.646	ENSRNOG00000025946	Igf2bp2	73	96.646	Rattus_norvegicus
ENSUAMG00000001610	-	94	98.476	ENSRBIG00000044700	IGF2BP2	78	98.476	Rhinopithecus_bieti
ENSUAMG00000001610	-	90	62.187	ENSRBIG00000004154	-	65	55.255	Rhinopithecus_bieti
ENSUAMG00000001610	-	94	69.512	ENSRBIG00000040002	-	54	69.512	Rhinopithecus_bieti
ENSUAMG00000001610	-	94	98.476	ENSRROG00000035074	IGF2BP2	75	98.476	Rhinopithecus_roxellana
ENSUAMG00000001610	-	95	54.706	ENSRROG00000017773	-	75	54.438	Rhinopithecus_roxellana
ENSUAMG00000001610	-	94	68.902	ENSRROG00000038130	-	52	68.902	Rhinopithecus_roxellana
ENSUAMG00000001610	-	94	98.476	ENSSBOG00000021548	IGF2BP2	78	98.476	Saimiri_boliviensis_boliviensis
ENSUAMG00000001610	-	93	83.434	ENSSHAG00000013152	IGF2BP2	78	83.434	Sarcophilus_harrisii
ENSUAMG00000001610	-	92	71.651	ENSSFOG00015007770	igf2bp2b	97	77.470	Scleropages_formosus
ENSUAMG00000001610	-	92	77.538	ENSSMAG00000010576	igf2bp2b	71	76.000	Scophthalmus_maximus
ENSUAMG00000001610	-	92	80.428	ENSSDUG00000017581	igf2bp2b	69	79.511	Seriola_dumerili
ENSUAMG00000001610	-	92	80.428	ENSSLDG00000007046	igf2bp2b	69	79.511	Seriola_lalandi_dorsalis
ENSUAMG00000001610	-	94	86.667	ENSSPUG00000016649	IGF2BP2	71	86.322	Sphenodon_punctatus
ENSUAMG00000001610	-	92	79.692	ENSSPAG00000000521	IGF2BP2	75	77.994	Stegastes_partitus
ENSUAMG00000001610	-	94	99.390	ENSSSCG00000011795	IGF2BP2	77	99.390	Sus_scrofa
ENSUAMG00000001610	-	94	85.410	ENSTGUG00000006719	-	78	85.410	Taeniopygia_guttata
ENSUAMG00000001610	-	92	75.077	ENSTRUG00000009908	igf2bp2b	67	74.154	Takifugu_rubripes
ENSUAMG00000001610	-	92	75.385	ENSTNIG00000016472	igf2bp2b	66	74.462	Tetraodon_nigroviridis
ENSUAMG00000001610	-	78	96.875	ENSTBEG00000006776	IGF2BP2	66	96.875	Tupaia_belangeri
ENSUAMG00000001610	-	94	98.476	ENSTTRG00000008241	IGF2BP2	70	98.476	Tursiops_truncatus
ENSUAMG00000001610	-	94	100.000	ENSUMAG00000004932	IGF2BP2	76	100.000	Ursus_maritimus
ENSUAMG00000001610	-	94	99.695	ENSVVUG00000012616	-	76	99.695	Vulpes_vulpes
ENSUAMG00000001610	-	92	74.925	ENSXCOG00000001170	igf2bp2b	65	78.616	Xiphophorus_couchianus
ENSUAMG00000001610	-	92	77.151	ENSXMAG00000015507	igf2bp2b	66	77.946	Xiphophorus_maculatus