EuRBPDB

Description

RBDs

Transcripts

Gene Model

PPI

Paralogs

Orthologs

Description

Ensembl ID: MGP_CAROLIEiJ_G0019016 (Gene tree)
Gene ID: 110309230
Gene Symbol: Polr3a
Alias: 9330175N20Rik|RPC1|RPC155
Full Name: RNA polymerase III subunit A
Gene Type: protein_coding
Species: Mus_caroli
Status: putative
Strand: Minus strand
Length: 39339 bases
Position: chr14:17159931-17199269
Accession: 110309230
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
MGP_CAROLIEiJ_P0035848	RNA_pol_Rpb1_5	PF04998.17	7.9e-89	1	1
MGP_CAROLIEiJ_P0035848	RNA_pol_Rpb1_4	PF05000.17	9.5e-32	1	1
MGP_CAROLIEiJ_P0035848	RNA_pol_Rpb1_2	PF00623.20	1.5e-74	1	1
MGP_CAROLIEiJ_P0035848	RNA_pol_Rpb1_3	PF04983.18	9e-42	1	1
MGP_CAROLIEiJ_P0035848	RNA_pol_Rpb1_1	PF04997.12	2.7e-96	1	1

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
MGP_CAROLIEiJ_T0035848	-	4685	XM_021181759	MGP_CAROLIEiJ_P0035848	1390 (aa)	XP_021037418	UPI000A319A9C

Gene Model

Click here to download MGP_CAROLIEiJ_G0019016's gene model file

Protein-Protein Interaction (PPI)

Clik here to download MGP_CAROLIEiJ_G0019016's network

Paralogs

Ensembl ID	Gene Symbol	Coverage	Identiy	Paralog	Gene Symbol	Coverage	Identiy
MGP_CAROLIEiJ_G0019016	Polr3a	91	37.939	MGP_CAROLIEiJ_G0016587	Polr2a	67	38.079
MGP_CAROLIEiJ_G0019016	Polr3a	77	32.551	MGP_CAROLIEiJ_G0028494	Polr1a	68	32.441

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.129	ENSG00000148606	POLR3A	100	98.129	Homo_sapiens
MGP_CAROLIEiJ_G0019016	Polr3a	62	86.179	ENSAPOG00000022727	polr3a	99	86.179	Acanthochromis_polyacanthus
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.990	ENSAMEG00000017801	POLR3A	100	97.990	Ailuropoda_melanoleuca
MGP_CAROLIEiJ_G0019016	Polr3a	93	90.439	ENSACIG00000024504	polr3a	97	90.439	Amphilophus_citrinellus
MGP_CAROLIEiJ_G0019016	Polr3a	100	87.050	ENSAOCG00000003949	-	100	87.050	Amphiprion_ocellaris
MGP_CAROLIEiJ_G0019016	Polr3a	100	90.360	ENSAOCG00000002071	polr3a	100	90.360	Amphiprion_ocellaris
MGP_CAROLIEiJ_G0019016	Polr3a	100	90.216	ENSAPEG00000001966	-	100	90.216	Amphiprion_percula
MGP_CAROLIEiJ_G0019016	Polr3a	100	90.360	ENSAPEG00000013351	polr3a	100	90.360	Amphiprion_percula
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.784	ENSATEG00000023277	polr3a	100	89.784	Anabas_testudineus
MGP_CAROLIEiJ_G0019016	Polr3a	99	95.855	ENSAPLG00000015878	POLR3A	99	95.855	Anas_platyrhynchos
MGP_CAROLIEiJ_G0019016	Polr3a	93	93.277	ENSACAG00000007985	POLR3A	99	93.277	Anolis_carolinensis
MGP_CAROLIEiJ_G0019016	Polr3a	100	95.334	ENSANAG00000028922	POLR3A	96	99.363	Aotus_nancymaae
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.353	ENSACLG00000005914	polr3a	100	89.353	Astatotilapia_calliptera
MGP_CAROLIEiJ_G0019016	Polr3a	100	87.482	ENSAMXG00000018852	polr3a	100	87.482	Astyanax_mexicanus
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.058	ENSBTAG00000013259	POLR3A	100	98.058	Bos_taurus
MGP_CAROLIEiJ_G0019016	Polr3a	99	56.797	WBGene00004411	rpc-1	99	56.797	Caenorhabditis_elegans
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.129	ENSCJAG00000001015	POLR3A	100	98.129	Callithrix_jacchus
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.273	ENSCAFG00000015546	POLR3A	100	98.273	Canis_familiaris
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.273	ENSCAFG00020014711	POLR3A	100	98.273	Canis_lupus_dingo
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.626	ENSCHIG00000000352	POLR3A	100	97.626	Capra_hircus
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.698	ENSTSYG00000006671	POLR3A	100	97.698	Carlito_syrichta
MGP_CAROLIEiJ_G0019016	Polr3a	99	97.382	ENSCAPG00000016031	POLR3A	100	97.382	Cavia_aperea
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.410	ENSCPOG00000022387	POLR3A	100	97.410	Cavia_porcellus
MGP_CAROLIEiJ_G0019016	Polr3a	100	95.971	ENSCCAG00000032995	POLR3A	100	95.971	Cebus_capucinus
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.914	ENSCATG00000035842	POLR3A	100	97.914	Cercocebus_atys
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.122	ENSCLAG00000009629	POLR3A	100	97.122	Chinchilla_lanigera
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.058	ENSCSAG00000009292	POLR3A	100	98.058	Chlorocebus_sabaeus
MGP_CAROLIEiJ_G0019016	Polr3a	85	82.686	ENSCHOG00000008368	POLR3A	100	82.686	Choloepus_hoffmanni
MGP_CAROLIEiJ_G0019016	Polr3a	99	94.888	ENSCPBG00000002079	POLR3A	99	94.888	Chrysemys_picta_bellii
MGP_CAROLIEiJ_G0019016	Polr3a	99	69.524	ENSCSAVG00000009925	-	99	69.524	Ciona_savignyi
MGP_CAROLIEiJ_G0019016	Polr3a	86	96.535	ENSCANG00000041071	POLR3A	100	96.535	Colobus_angolensis_palliatus
MGP_CAROLIEiJ_G0019016	Polr3a	78	99.462	ENSCGRG00001015599	-	92	99.462	Cricetulus_griseus_chok1gshd
MGP_CAROLIEiJ_G0019016	Polr3a	88	99.120	ENSCGRG00000001019	Polr3a	100	99.120	Cricetulus_griseus_crigri
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.928	ENSCSEG00000016653	polr3a	100	89.928	Cynoglossus_semilaevis
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.424	ENSCVAG00000023049	polr3a	100	89.424	Cyprinodon_variegatus
MGP_CAROLIEiJ_G0019016	Polr3a	99	90.425	ENSDARG00000102569	polr3a	99	90.425	Danio_rerio
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.410	ENSDNOG00000010194	POLR3A	100	97.410	Dasypus_novemcinctus
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.345	ENSDORG00000000184	Polr3a	100	98.345	Dipodomys_ordii
MGP_CAROLIEiJ_G0019016	Polr3a	99	57.236	FBgn0030687	CG17209	99	57.236	Drosophila_melanogaster
MGP_CAROLIEiJ_G0019016	Polr3a	88	97.166	ENSETEG00000007477	POLR3A	89	97.166	Echinops_telfairi
MGP_CAROLIEiJ_G0019016	Polr3a	59	76.698	ENSEBUG00000013761	polr3a	98	76.698	Eptatretus_burgeri
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.345	ENSEASG00005023053	POLR3A	100	98.345	Equus_asinus_asinus
MGP_CAROLIEiJ_G0019016	Polr3a	97	98.212	ENSECAG00000019416	POLR3A	100	98.053	Equus_caballus
MGP_CAROLIEiJ_G0019016	Polr3a	96	93.139	ENSEEUG00000001774	POLR3A	100	93.139	Erinaceus_europaeus
MGP_CAROLIEiJ_G0019016	Polr3a	100	87.940	ENSELUG00000014016	polr3a	100	87.940	Esox_lucius
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.489	ENSFCAG00000000843	POLR3A	100	98.489	Felis_catus
MGP_CAROLIEiJ_G0019016	Polr3a	100	95.615	ENSFALG00000006913	POLR3A	100	95.615	Ficedula_albicollis
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.410	ENSFDAG00000007813	POLR3A	100	97.410	Fukomys_damarensis
MGP_CAROLIEiJ_G0019016	Polr3a	95	89.737	ENSFHEG00000011922	polr3a	99	89.737	Fundulus_heteroclitus
MGP_CAROLIEiJ_G0019016	Polr3a	100	84.713	ENSGMOG00000000291	polr3a	99	84.713	Gadus_morhua
MGP_CAROLIEiJ_G0019016	Polr3a	100	95.971	ENSGALG00000004947	POLR3A	100	95.971	Gallus_gallus
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.496	ENSGAFG00000005112	polr3a	100	89.496	Gambusia_affinis
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.209	ENSGACG00000002304	polr3a	100	89.209	Gasterosteus_aculeatus
MGP_CAROLIEiJ_G0019016	Polr3a	95	95.155	ENSGAGG00000010386	POLR3A	98	95.155	Gopherus_agassizii
MGP_CAROLIEiJ_G0019016	Polr3a	99	97.038	ENSGGOG00000027729	POLR3A	100	98.213	Gorilla_gorilla
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.353	ENSHBUG00000018098	polr3a	100	89.353	Haplochromis_burtoni
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.554	ENSHGLG00000017899	POLR3A	100	97.554	Heterocephalus_glaber_female
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.554	ENSHGLG00100008483	POLR3A	100	97.554	Heterocephalus_glaber_male
MGP_CAROLIEiJ_G0019016	Polr3a	100	88.993	ENSHCOG00000001075	polr3a	100	88.993	Hippocampus_comes
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.345	ENSSTOG00000012983	POLR3A	100	98.345	Ictidomys_tridecemlineatus
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.784	ENSKMAG00000015290	polr3a	100	89.784	Kryptolebias_marmoratus
MGP_CAROLIEiJ_G0019016	Polr3a	99	88.769	ENSLBEG00000017214	polr3a	99	88.769	Labrus_bergylta
MGP_CAROLIEiJ_G0019016	Polr3a	97	89.740	ENSLACG00000018557	POLR3A	99	89.740	Latimeria_chalumnae
MGP_CAROLIEiJ_G0019016	Polr3a	100	91.744	ENSLOCG00000004942	polr3a	100	91.744	Lepisosteus_oculatus
MGP_CAROLIEiJ_G0019016	Polr3a	100	96.334	ENSLAFG00000016719	POLR3A	100	96.334	Loxodonta_africana
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.986	ENSMFAG00000032218	POLR3A	100	97.986	Macaca_fascicularis
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.986	ENSMMUG00000005514	POLR3A	100	97.986	Macaca_mulatta
MGP_CAROLIEiJ_G0019016	Polr3a	99	90.708	ENSMNEG00000035163	POLR3A	99	97.865	Macaca_nemestrina
MGP_CAROLIEiJ_G0019016	Polr3a	100	93.391	ENSMLEG00000030014	POLR3A	98	98.734	Mandrillus_leucophaeus
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.640	ENSMAMG00000017497	polr3a	100	89.640	Mastacembelus_armatus
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.353	ENSMZEG00005000576	polr3a	100	89.353	Maylandia_zebra
MGP_CAROLIEiJ_G0019016	Polr3a	100	77.557	ENSMGAG00000005713	-	100	77.557	Meleagris_gallopavo
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.194	ENSMAUG00000016247	Polr3a	100	97.194	Mesocricetus_auratus
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.345	ENSMICG00000005937	POLR3A	100	98.436	Microcebus_murinus
MGP_CAROLIEiJ_G0019016	Polr3a	100	99.209	ENSMOCG00000012185	Polr3a	100	99.209	Microtus_ochrogaster
MGP_CAROLIEiJ_G0019016	Polr3a	100	88.993	ENSMMOG00000013822	polr3a	100	88.993	Mola_mola
MGP_CAROLIEiJ_G0019016	Polr3a	100	96.978	ENSMODG00000001036	POLR3A	100	96.978	Monodelphis_domestica
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.137	ENSMALG00000001249	polr3a	100	89.137	Monopterus_albus
MGP_CAROLIEiJ_G0019016	Polr3a	100	99.856	ENSMUSG00000025280	Polr3a	100	99.856	Mus_musculus
MGP_CAROLIEiJ_G0019016	Polr3a	100	99.856	MGP_PahariEiJ_G0030029	Polr3a	100	99.856	Mus_pahari
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.410	MGP_SPRETEiJ_G0019894	Polr3a	100	97.410	Mus_spretus
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.575	ENSMPUG00000003790	POLR3A	100	97.575	Mustela_putorius_furo
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.266	ENSMLUG00000011522	POLR3A	100	97.266	Myotis_lucifugus
MGP_CAROLIEiJ_G0019016	Polr3a	100	96.835	ENSNGAG00000015863	Polr3a	100	96.835	Nannospalax_galili
MGP_CAROLIEiJ_G0019016	Polr3a	100	88.058	ENSNBRG00000014262	polr3a	100	88.058	Neolamprologus_brichardi
MGP_CAROLIEiJ_G0019016	Polr3a	88	92.208	ENSNLEG00000010501	POLR3A	92	92.208	Nomascus_leucogenys
MGP_CAROLIEiJ_G0019016	Polr3a	99	84.145	ENSMEUG00000012446	POLR3A	100	84.145	Notamacropus_eugenii
MGP_CAROLIEiJ_G0019016	Polr3a	100	92.734	ENSOPRG00000001880	-	100	92.734	Ochotona_princeps
MGP_CAROLIEiJ_G0019016	Polr3a	100	95.899	ENSODEG00000015146	POLR3A	100	95.899	Octodon_degus
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.784	ENSONIG00000016129	polr3a	100	89.784	Oreochromis_niloticus
MGP_CAROLIEiJ_G0019016	Polr3a	98	96.275	ENSOANG00000007013	POLR3A	100	96.275	Ornithorhynchus_anatinus
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.698	ENSOCUG00000012450	POLR3A	100	97.698	Oryctolagus_cuniculus
MGP_CAROLIEiJ_G0019016	Polr3a	100	90.000	ENSORLG00000008958	polr3a	100	90.000	Oryzias_latipes
MGP_CAROLIEiJ_G0019016	Polr3a	100	87.770	ENSORLG00020007525	polr3a	98	90.313	Oryzias_latipes_hni
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.856	ENSORLG00015006633	polr3a	100	89.856	Oryzias_latipes_hsok
MGP_CAROLIEiJ_G0019016	Polr3a	100	88.921	ENSOMEG00000012865	polr3a	100	88.921	Oryzias_melastigma
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.058	ENSOGAG00000010995	POLR3A	100	98.058	Otolemur_garnettii
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.629	ENSOARG00000009656	POLR3A	100	97.629	Ovis_aries
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.489	ENSPPRG00000002128	POLR3A	100	98.489	Panthera_pardus
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.489	ENSPTIG00000020916	POLR3A	100	98.489	Panthera_tigris_altaica
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.058	ENSPTRG00000002664	POLR3A	100	98.058	Pan_troglodytes
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.914	ENSPANG00000022508	POLR3A	100	97.914	Papio_anubis
MGP_CAROLIEiJ_G0019016	Polr3a	100	91.439	ENSPKIG00000008140	polr3a	100	91.439	Paramormyrops_kingsleyae
MGP_CAROLIEiJ_G0019016	Polr3a	98	94.960	ENSPSIG00000002092	POLR3A	100	94.960	Pelodiscus_sinensis
MGP_CAROLIEiJ_G0019016	Polr3a	100	99.209	ENSPEMG00000006768	Polr3a	100	99.209	Peromyscus_maniculatus_bairdii
MGP_CAROLIEiJ_G0019016	Polr3a	100	96.835	ENSPCIG00000015842	POLR3A	100	96.835	Phascolarctos_cinereus
MGP_CAROLIEiJ_G0019016	Polr3a	54	90.872	ENSPFOG00000006458	polr3a	100	90.872	Poecilia_formosa
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.784	ENSPLAG00000000997	polr3a	100	89.784	Poecilia_latipinna
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.712	ENSPMEG00000006821	polr3a	100	89.712	Poecilia_mexicana
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.640	ENSPREG00000002385	polr3a	100	89.640	Poecilia_reticulata
MGP_CAROLIEiJ_G0019016	Polr3a	93	97.757	ENSPPYG00000002305	POLR3A	99	97.757	Pongo_abelii
MGP_CAROLIEiJ_G0019016	Polr3a	84	85.115	ENSPCAG00000005690	POLR3A	87	85.115	Procavia_capensis
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.986	ENSPCOG00000028311	POLR3A	100	97.986	Propithecus_coquereli
MGP_CAROLIEiJ_G0019016	Polr3a	100	96.906	ENSPVAG00000015553	POLR3A	100	96.906	Pteropus_vampyrus
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.353	ENSPNYG00000020987	polr3a	100	89.353	Pundamilia_nyererei
MGP_CAROLIEiJ_G0019016	Polr3a	99	86.393	ENSPNAG00000006629	polr3a	99	86.393	Pygocentrus_nattereri
MGP_CAROLIEiJ_G0019016	Polr3a	100	99.496	ENSRNOG00000010008	Polr3a	100	99.496	Rattus_norvegicus
MGP_CAROLIEiJ_G0019016	Polr3a	100	93.597	ENSRBIG00000034967	POLR3A	98	98.734	Rhinopithecus_bieti
MGP_CAROLIEiJ_G0019016	Polr3a	100	97.842	ENSRROG00000025527	POLR3A	100	97.842	Rhinopithecus_roxellana
MGP_CAROLIEiJ_G0019016	Polr3a	99	50.276	YOR116C	RPO31	99	50.276	Saccharomyces_cerevisiae
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.201	ENSSBOG00000022199	POLR3A	100	98.201	Saimiri_boliviensis_boliviensis
MGP_CAROLIEiJ_G0019016	Polr3a	100	96.691	ENSSHAG00000007588	POLR3A	94	96.691	Sarcophilus_harrisii
MGP_CAROLIEiJ_G0019016	Polr3a	100	91.655	ENSSFOG00015014357	polr3a	100	91.655	Scleropages_formosus
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.424	ENSSMAG00000001780	polr3a	100	89.424	Scophthalmus_maximus
MGP_CAROLIEiJ_G0019016	Polr3a	100	90.072	ENSSDUG00000019840	polr3a	100	90.072	Seriola_dumerili
MGP_CAROLIEiJ_G0019016	Polr3a	100	90.072	ENSSLDG00000023908	polr3a	100	90.072	Seriola_lalandi_dorsalis
MGP_CAROLIEiJ_G0019016	Polr3a	69	98.746	ENSSARG00000009916	POLR3A	89	98.746	Sorex_araneus
MGP_CAROLIEiJ_G0019016	Polr3a	100	93.237	ENSSPUG00000013209	POLR3A	100	93.237	Sphenodon_punctatus
MGP_CAROLIEiJ_G0019016	Polr3a	100	90.216	ENSSPAG00000017712	polr3a	100	90.216	Stegastes_partitus
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.058	ENSSSCG00000010327	POLR3A	100	98.058	Sus_scrofa
MGP_CAROLIEiJ_G0019016	Polr3a	100	95.540	ENSTGUG00000006423	-	100	95.540	Taeniopygia_guttata
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.137	ENSTRUG00000013488	polr3a	100	89.137	Takifugu_rubripes
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.368	ENSTNIG00000013151	polr3a	100	89.368	Tetraodon_nigroviridis
MGP_CAROLIEiJ_G0019016	Polr3a	71	96.809	ENSTBEG00000015288	POLR3A	75	96.809	Tupaia_belangeri
MGP_CAROLIEiJ_G0019016	Polr3a	96	98.555	ENSTTRG00000007817	POLR3A	100	98.555	Tursiops_truncatus
MGP_CAROLIEiJ_G0019016	Polr3a	100	95.899	ENSUAMG00000011283	POLR3A	100	95.899	Ursus_americanus
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.273	ENSUMAG00000013913	POLR3A	100	98.273	Ursus_maritimus
MGP_CAROLIEiJ_G0019016	Polr3a	99	89.891	ENSVPAG00000003285	POLR3A	100	89.891	Vicugna_pacos
MGP_CAROLIEiJ_G0019016	Polr3a	100	98.201	ENSVVUG00000025523	POLR3A	100	98.201	Vulpes_vulpes
MGP_CAROLIEiJ_G0019016	Polr3a	86	92.314	ENSXETG00000018101	polr3a	100	92.314	Xenopus_tropicalis
MGP_CAROLIEiJ_G0019016	Polr3a	100	89.496	ENSXMAG00000015213	polr3a	100	89.496	Xiphophorus_maculatus