EuRBPDB

Description

RBDs

Transcripts

Gene Model

PPI

Orthologs

Description

Ensembl ID: MGP_CAROLIEiJ_G0022733 (Gene tree)
Gene ID: 110285379
Gene Symbol: Mrpl16
Alias: 2310039D06Rik
Full Name: mitochondrial ribosomal protein L16
Gene Type: protein_coding
Species: Mus_caroli
Status: putative
Strand: Plus strand
Length: 4509 bases
Position: chr19:9234451-9238959
Accession: 110285379
RBP type: canonical RBP
Summary: -

RNA binding domains(RBDs)

Protein ID	Domain	Pfam ID	E-value	Domain number	Total number
MGP_CAROLIEiJ_P0048244	Ribosomal_L16	PF00252.18	1.4e-29	1	1

Transcripts

Transcript ID	Name	Length	RefSeq ID	Protein ID	Length	RefSeq ID	UniportKB ID
MGP_CAROLIEiJ_T0048244	-	1122	-	MGP_CAROLIEiJ_P0048244	251 (aa)	XP_021007146	UPI000A31AA1C

Gene Model

Click here to download MGP_CAROLIEiJ_G0022733's gene model file

Protein-Protein Interaction (PPI)

Clik here to download MGP_CAROLIEiJ_G0022733's network

Orthologs

Ensembl ID	Gene Symbol	Coverage	Identiy	Ortholog	Gene Symbol	Coverage	Identiy	Species
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.861	ENSG00000166902	MRPL16	100	89.535	Homo_sapiens
MGP_CAROLIEiJ_G0022733	Mrpl16	89	57.589	ENSAPOG00000003818	mrpl16	87	57.589	Acanthochromis_polyacanthus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.861	ENSAMEG00000014289	MRPL16	100	84.861	Ailuropoda_melanoleuca
MGP_CAROLIEiJ_G0022733	Mrpl16	89	59.375	ENSACIG00000022204	mrpl16	87	59.375	Amphilophus_citrinellus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	59.375	ENSAOCG00000002647	mrpl16	87	59.375	Amphiprion_ocellaris
MGP_CAROLIEiJ_G0022733	Mrpl16	89	59.375	ENSAPEG00000020339	mrpl16	87	59.375	Amphiprion_percula
MGP_CAROLIEiJ_G0022733	Mrpl16	89	62.946	ENSATEG00000012565	mrpl16	87	62.946	Anabas_testudineus
MGP_CAROLIEiJ_G0022733	Mrpl16	99	66.932	ENSACAG00000009952	MRPL16	99	66.932	Anolis_carolinensis
MGP_CAROLIEiJ_G0022733	Mrpl16	100	85.259	ENSANAG00000029141	MRPL16	100	85.259	Aotus_nancymaae
MGP_CAROLIEiJ_G0022733	Mrpl16	87	58.904	ENSACLG00000024282	mrpl16	95	58.904	Astatotilapia_calliptera
MGP_CAROLIEiJ_G0022733	Mrpl16	89	62.054	ENSAMXG00000014688	mrpl16	92	62.054	Astyanax_mexicanus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.861	ENSBTAG00000021188	MRPL16	100	84.861	Bos_taurus
MGP_CAROLIEiJ_G0022733	Mrpl16	95	30.000	WBGene00011412	mrpl-16	98	30.000	Caenorhabditis_elegans
MGP_CAROLIEiJ_G0022733	Mrpl16	100	85.657	ENSCJAG00000016306	MRPL16	100	85.657	Callithrix_jacchus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.861	ENSCAFG00000031711	-	100	84.861	Canis_familiaris
MGP_CAROLIEiJ_G0022733	Mrpl16	99	82.731	ENSCAFG00000029686	-	99	82.731	Canis_familiaris
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.861	ENSCAFG00020000230	MRPL16	100	84.861	Canis_lupus_dingo
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.861	ENSCHIG00000012712	MRPL16	100	84.861	Capra_hircus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	86.454	ENSTSYG00000013336	MRPL16	100	86.454	Carlito_syrichta
MGP_CAROLIEiJ_G0022733	Mrpl16	100	82.869	ENSCPOG00000005035	MRPL16	100	82.869	Cavia_porcellus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.861	ENSCCAG00000035372	MRPL16	100	84.861	Cebus_capucinus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.462	ENSCATG00000035790	MRPL16	100	84.462	Cercocebus_atys
MGP_CAROLIEiJ_G0022733	Mrpl16	89	82.511	ENSCLAG00000003811	MRPL16	100	78.884	Chinchilla_lanigera
MGP_CAROLIEiJ_G0022733	Mrpl16	100	85.657	ENSCSAG00000006107	MRPL16	100	85.657	Chlorocebus_sabaeus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	68.526	ENSCHOG00000009976	MRPL16	100	68.526	Choloepus_hoffmanni
MGP_CAROLIEiJ_G0022733	Mrpl16	100	67.331	ENSCPBG00000009578	MRPL16	100	67.331	Chrysemys_picta_bellii
MGP_CAROLIEiJ_G0022733	Mrpl16	62	39.103	ENSCING00000020946	-	85	39.103	Ciona_intestinalis
MGP_CAROLIEiJ_G0022733	Mrpl16	100	83.267	ENSCANG00000027375	MRPL16	100	83.267	Colobus_angolensis_palliatus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	88.845	ENSCGRG00001000785	Mrpl16	100	88.845	Cricetulus_griseus_chok1gshd
MGP_CAROLIEiJ_G0022733	Mrpl16	84	89.048	ENSCGRG00000015073	Mrpl16	100	89.048	Cricetulus_griseus_crigri
MGP_CAROLIEiJ_G0022733	Mrpl16	89	56.696	ENSCSEG00000006169	mrpl16	87	56.696	Cynoglossus_semilaevis
MGP_CAROLIEiJ_G0022733	Mrpl16	90	60.177	ENSCVAG00000013739	mrpl16	88	60.177	Cyprinodon_variegatus
MGP_CAROLIEiJ_G0022733	Mrpl16	90	60.177	ENSCVAG00000014885	mrpl16	88	60.177	Cyprinodon_variegatus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	60.268	ENSDARG00000075325	mrpl16	97	61.395	Danio_rerio
MGP_CAROLIEiJ_G0022733	Mrpl16	80	59.701	ENSDNOG00000041871	-	99	59.701	Dasypus_novemcinctus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	79.283	ENSDNOG00000034262	-	100	79.283	Dasypus_novemcinctus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.861	ENSDORG00000024946	Mrpl16	100	84.861	Dipodomys_ordii
MGP_CAROLIEiJ_G0022733	Mrpl16	90	46.930	FBgn0023519	mRpL16	94	46.930	Drosophila_melanogaster
MGP_CAROLIEiJ_G0022733	Mrpl16	100	65.476	ENSETEG00000011252	MRPL16	100	65.476	Echinops_telfairi
MGP_CAROLIEiJ_G0022733	Mrpl16	89	61.607	ENSEBUG00000001257	mrpl16	87	61.607	Eptatretus_burgeri
MGP_CAROLIEiJ_G0022733	Mrpl16	100	86.454	ENSEASG00005007994	MRPL16	100	86.454	Equus_asinus_asinus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	86.454	ENSECAG00000014926	MRPL16	100	86.454	Equus_caballus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	81.275	ENSEEUG00000011076	MRPL16	100	81.275	Erinaceus_europaeus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	61.161	ENSELUG00000002608	mrpl16	86	61.161	Esox_lucius
MGP_CAROLIEiJ_G0022733	Mrpl16	100	83.665	ENSFCAG00000007540	MRPL16	100	83.665	Felis_catus
MGP_CAROLIEiJ_G0022733	Mrpl16	84	69.524	ENSFALG00000012123	MRPL16	99	69.524	Ficedula_albicollis
MGP_CAROLIEiJ_G0022733	Mrpl16	92	87.500	ENSFDAG00000012285	MRPL16	100	87.500	Fukomys_damarensis
MGP_CAROLIEiJ_G0022733	Mrpl16	89	62.500	ENSFHEG00000015359	mrpl16	85	62.500	Fundulus_heteroclitus
MGP_CAROLIEiJ_G0022733	Mrpl16	65	62.346	ENSGMOG00000008546	mrpl16	85	62.346	Gadus_morhua
MGP_CAROLIEiJ_G0022733	Mrpl16	79	70.352	ENSGALG00000029419	MRPL16	100	70.352	Gallus_gallus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	59.375	ENSGAFG00000012264	mrpl16	87	59.375	Gambusia_affinis
MGP_CAROLIEiJ_G0022733	Mrpl16	89	58.482	ENSGACG00000015363	mrpl16	87	58.482	Gasterosteus_aculeatus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	66.534	ENSGAGG00000023594	MRPL16	100	66.534	Gopherus_agassizii
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.462	ENSGGOG00000015703	MRPL16	100	84.462	Gorilla_gorilla
MGP_CAROLIEiJ_G0022733	Mrpl16	89	59.821	ENSHBUG00000001596	mrpl16	87	59.821	Haplochromis_burtoni
MGP_CAROLIEiJ_G0022733	Mrpl16	100	82.072	ENSHGLG00000010777	MRPL16	100	82.072	Heterocephalus_glaber_female
MGP_CAROLIEiJ_G0022733	Mrpl16	100	82.072	ENSHGLG00100010477	MRPL16	100	82.072	Heterocephalus_glaber_male
MGP_CAROLIEiJ_G0022733	Mrpl16	89	61.161	ENSHCOG00000003499	mrpl16	91	61.792	Hippocampus_comes
MGP_CAROLIEiJ_G0022733	Mrpl16	89	64.286	ENSIPUG00000009966	mrpl16	87	64.286	Ictalurus_punctatus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	85.657	ENSSTOG00000021462	MRPL16	100	85.657	Ictidomys_tridecemlineatus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	85.259	ENSJJAG00000018928	Mrpl16	100	85.259	Jaculus_jaculus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	56.696	ENSKMAG00000022035	mrpl16	89	56.696	Kryptolebias_marmoratus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	61.161	ENSLBEG00000013193	mrpl16	87	61.161	Labrus_bergylta
MGP_CAROLIEiJ_G0022733	Mrpl16	89	68.750	ENSLACG00000002060	MRPL16	89	68.750	Latimeria_chalumnae
MGP_CAROLIEiJ_G0022733	Mrpl16	89	64.732	ENSLOCG00000010979	mrpl16	88	64.732	Lepisosteus_oculatus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	80.876	ENSLAFG00000001490	MRPL16	100	80.876	Loxodonta_africana
MGP_CAROLIEiJ_G0022733	Mrpl16	100	85.259	ENSMFAG00000002379	MRPL16	100	85.259	Macaca_fascicularis
MGP_CAROLIEiJ_G0022733	Mrpl16	100	85.259	ENSMMUG00000022321	MRPL16	100	85.259	Macaca_mulatta
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.064	ENSMNEG00000040325	MRPL16	100	84.064	Macaca_nemestrina
MGP_CAROLIEiJ_G0022733	Mrpl16	100	77.291	ENSMLEG00000033890	MRPL16	100	77.291	Mandrillus_leucophaeus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	61.607	ENSMAMG00000003085	mrpl16	93	62.085	Mastacembelus_armatus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	59.821	ENSMZEG00005000716	mrpl16	87	59.821	Maylandia_zebra
MGP_CAROLIEiJ_G0022733	Mrpl16	99	62.598	ENSMGAG00000003328	MRPL16	99	62.598	Meleagris_gallopavo
MGP_CAROLIEiJ_G0022733	Mrpl16	100	90.837	ENSMAUG00000011181	Mrpl16	100	90.837	Mesocricetus_auratus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.861	ENSMICG00000011027	MRPL16	100	84.861	Microcebus_murinus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	90.837	ENSMOCG00000003604	Mrpl16	100	90.837	Microtus_ochrogaster
MGP_CAROLIEiJ_G0022733	Mrpl16	89	59.821	ENSMMOG00000018112	mrpl16	86	59.821	Mola_mola
MGP_CAROLIEiJ_G0022733	Mrpl16	100	74.104	ENSMODG00000020205	MRPL16	100	74.104	Monodelphis_domestica
MGP_CAROLIEiJ_G0022733	Mrpl16	89	60.714	ENSMALG00000016263	mrpl16	89	60.714	Monopterus_albus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	98.406	ENSMUSG00000024683	Mrpl16	100	98.406	Mus_musculus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	97.211	MGP_PahariEiJ_G0014227	Mrpl16	100	97.211	Mus_pahari
MGP_CAROLIEiJ_G0022733	Mrpl16	100	98.008	MGP_SPRETEiJ_G0023648	Mrpl16	100	98.008	Mus_spretus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.861	ENSMPUG00000014583	MRPL16	100	84.861	Mustela_putorius_furo
MGP_CAROLIEiJ_G0022733	Mrpl16	100	85.259	ENSMLUG00000011224	-	100	85.259	Myotis_lucifugus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	74.502	ENSMLUG00000027025	-	100	74.502	Myotis_lucifugus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	85.259	ENSNGAG00000010636	Mrpl16	100	85.259	Nannospalax_galili
MGP_CAROLIEiJ_G0022733	Mrpl16	89	59.375	ENSNBRG00000017868	mrpl16	87	59.375	Neolamprologus_brichardi
MGP_CAROLIEiJ_G0022733	Mrpl16	100	85.657	ENSNLEG00000013137	MRPL16	100	85.657	Nomascus_leucogenys
MGP_CAROLIEiJ_G0022733	Mrpl16	84	79.048	ENSMEUG00000016437	MRPL16	100	79.048	Notamacropus_eugenii
MGP_CAROLIEiJ_G0022733	Mrpl16	100	85.317	ENSOPRG00000014638	-	100	85.317	Ochotona_princeps
MGP_CAROLIEiJ_G0022733	Mrpl16	100	80.478	ENSODEG00000001664	MRPL16	100	80.478	Octodon_degus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	59.375	ENSONIG00000003122	mrpl16	87	59.375	Oreochromis_niloticus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	86.056	ENSOCUG00000013390	MRPL16	100	86.056	Oryctolagus_cuniculus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	58.929	ENSORLG00000026229	mrpl16	87	58.929	Oryzias_latipes
MGP_CAROLIEiJ_G0022733	Mrpl16	89	58.929	ENSORLG00020016055	mrpl16	65	58.929	Oryzias_latipes_hni
MGP_CAROLIEiJ_G0022733	Mrpl16	89	58.929	ENSORLG00015014562	mrpl16	82	58.929	Oryzias_latipes_hsok
MGP_CAROLIEiJ_G0022733	Mrpl16	89	59.821	ENSOMEG00000016848	mrpl16	87	59.821	Oryzias_melastigma
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.861	ENSOGAG00000001486	MRPL16	100	84.861	Otolemur_garnettii
MGP_CAROLIEiJ_G0022733	Mrpl16	100	80.876	ENSOARG00000012266	MRPL16	100	80.876	Ovis_aries
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.861	ENSPPAG00000038990	MRPL16	100	84.861	Pan_paniscus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	83.665	ENSPPRG00000013233	MRPL16	100	83.665	Panthera_pardus
MGP_CAROLIEiJ_G0022733	Mrpl16	92	85.345	ENSPTIG00000014818	MRPL16	92	85.345	Panthera_tigris_altaica
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.861	ENSPTRG00000003700	MRPL16	100	84.861	Pan_troglodytes
MGP_CAROLIEiJ_G0022733	Mrpl16	100	85.259	ENSPANG00000010272	MRPL16	100	85.259	Papio_anubis
MGP_CAROLIEiJ_G0022733	Mrpl16	93	61.803	ENSPKIG00000010334	mrpl16	91	61.803	Paramormyrops_kingsleyae
MGP_CAROLIEiJ_G0022733	Mrpl16	100	67.331	ENSPSIG00000016182	MRPL16	100	67.331	Pelodiscus_sinensis
MGP_CAROLIEiJ_G0022733	Mrpl16	89	62.946	ENSPMGG00000001444	mrpl16	88	62.946	Periophthalmus_magnuspinnatus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	90.837	ENSPEMG00000001887	Mrpl16	100	90.837	Peromyscus_maniculatus_bairdii
MGP_CAROLIEiJ_G0022733	Mrpl16	94	58.264	ENSPMAG00000001603	mrpl16	92	58.264	Petromyzon_marinus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	77.291	ENSPCIG00000009443	MRPL16	100	77.291	Phascolarctos_cinereus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	61.161	ENSPFOG00000009144	mrpl16	87	61.161	Poecilia_formosa
MGP_CAROLIEiJ_G0022733	Mrpl16	89	61.161	ENSPLAG00000010811	mrpl16	87	61.161	Poecilia_latipinna
MGP_CAROLIEiJ_G0022733	Mrpl16	89	61.161	ENSPMEG00000023095	mrpl16	87	61.161	Poecilia_mexicana
MGP_CAROLIEiJ_G0022733	Mrpl16	89	60.714	ENSPREG00000021826	mrpl16	95	60.714	Poecilia_reticulata
MGP_CAROLIEiJ_G0022733	Mrpl16	100	83.665	ENSPPYG00000003227	MRPL16	100	83.665	Pongo_abelii
MGP_CAROLIEiJ_G0022733	Mrpl16	100	80.080	ENSPCAG00000008055	MRPL16	100	80.080	Procavia_capensis
MGP_CAROLIEiJ_G0022733	Mrpl16	100	85.259	ENSPCOG00000025205	MRPL16	100	85.259	Propithecus_coquereli
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.462	ENSPVAG00000012250	MRPL16	100	84.462	Pteropus_vampyrus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	59.821	ENSPNYG00000003647	mrpl16	87	59.821	Pundamilia_nyererei
MGP_CAROLIEiJ_G0022733	Mrpl16	89	59.821	ENSPNAG00000010473	mrpl16	88	59.821	Pygocentrus_nattereri
MGP_CAROLIEiJ_G0022733	Mrpl16	100	96.016	ENSRNOG00000021005	Mrpl16	100	96.016	Rattus_norvegicus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.462	ENSRBIG00000018289	MRPL16	100	84.462	Rhinopithecus_bieti
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.462	ENSRROG00000040273	MRPL16	100	84.462	Rhinopithecus_roxellana
MGP_CAROLIEiJ_G0022733	Mrpl16	93	84.549	ENSSBOG00000005353	MRPL16	94	84.834	Saimiri_boliviensis_boliviensis
MGP_CAROLIEiJ_G0022733	Mrpl16	100	74.502	ENSSHAG00000006665	MRPL16	100	74.502	Sarcophilus_harrisii
MGP_CAROLIEiJ_G0022733	Mrpl16	89	62.500	ENSSFOG00015008725	mrpl16	89	64.762	Scleropages_formosus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	58.036	ENSSMAG00000021324	mrpl16	92	58.036	Scophthalmus_maximus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	58.482	ENSSDUG00000006482	mrpl16	87	58.482	Seriola_dumerili
MGP_CAROLIEiJ_G0022733	Mrpl16	89	58.482	ENSSLDG00000000994	mrpl16	87	58.482	Seriola_lalandi_dorsalis
MGP_CAROLIEiJ_G0022733	Mrpl16	84	86.667	ENSSARG00000004666	MRPL16	100	86.667	Sorex_araneus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	66.534	ENSSPUG00000018047	MRPL16	100	66.534	Sphenodon_punctatus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	58.036	ENSSPAG00000017237	mrpl16	87	58.036	Stegastes_partitus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	83.267	ENSSSCG00000013118	MRPL16	100	83.267	Sus_scrofa
MGP_CAROLIEiJ_G0022733	Mrpl16	59	65.541	ENSTGUG00000006277	MRPL16	100	65.541	Taeniopygia_guttata
MGP_CAROLIEiJ_G0022733	Mrpl16	89	53.571	ENSTRUG00000017231	mrpl16	86	53.571	Takifugu_rubripes
MGP_CAROLIEiJ_G0022733	Mrpl16	89	60.444	ENSTNIG00000009438	mrpl16	88	60.444	Tetraodon_nigroviridis
MGP_CAROLIEiJ_G0022733	Mrpl16	100	82.072	ENSTBEG00000007959	MRPL16	100	82.072	Tupaia_belangeri
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.064	ENSTTRG00000009428	MRPL16	100	84.064	Tursiops_truncatus
MGP_CAROLIEiJ_G0022733	Mrpl16	100	84.462	ENSUAMG00000009824	MRPL16	99	86.301	Ursus_americanus
MGP_CAROLIEiJ_G0022733	Mrpl16	92	85.776	ENSUMAG00000018814	MRPL16	99	86.758	Ursus_maritimus
MGP_CAROLIEiJ_G0022733	Mrpl16	92	86.522	ENSVVUG00000026682	MRPL16	95	86.522	Vulpes_vulpes
MGP_CAROLIEiJ_G0022733	Mrpl16	94	58.824	ENSXETG00000024210	mrpl16	94	58.824	Xenopus_tropicalis
MGP_CAROLIEiJ_G0022733	Mrpl16	89	60.714	ENSXCOG00000002880	mrpl16	90	60.714	Xiphophorus_couchianus
MGP_CAROLIEiJ_G0022733	Mrpl16	89	60.714	ENSXMAG00000019187	mrpl16	87	60.714	Xiphophorus_maculatus