Protein kinase%3B along with its paralog%2C ALK2%2C required for proper spindle positioning and nuclear segregation following mitotic arrest%2C proper organization of cell polarity factors in mitosis%2C proper localization of formins and polarity factors%2C and survival in cells that activate spindle assembly checkpoint%3B phosphorylated in response to DNA damage%3B ALK1 has a paralog%2C ALK2%2C that arose from the whole genome duplication%3B similar to mammalian haspins